Helicosina valga Kuwahara & Marshall & Yau 2024, sp. nov.

Helicosina valga Marshall sp. nov. (Figs. 2E–F, 3C, 6, 8) Description. Body length: Males 1.9–2.1 mm, females 2.0– 2.2 mm. General (Fig. 2E): Male fore and hind femora strongly thickened (length 2.9–3.0 × and 3.5–3.7 × width, respectively). Male abdomen (Fig. 6): S5 broad, gently curved, broadly desclerotized posteromedially, and densely setose laterally. Posterior opening of epandrium oval (medial height 1.3–1.4 × medial width); cercus separated medially, forming a weakly-developed subanal plate. Surstylus strongly sinuate, basal half broad and subtriangular with the apex curved posteriorly, apical half strongly bent downwards and constricted with several anterior setae. Postgonite small, broad, and apically tapered with a small subapical patch of microtomentum. Phallapodeme elongate and well sclerotized. Basiphallus stout, slightly curved, with a stout, apically pointed epiphallus; distiphallus stout, largely membranous but including a trough-shaped basal sclerite with upper surface bent inwards to form a “roof”, a shieldshaped upper sclerite, and paired elongate lower-lateral sclerites; lateral membrane densely microtrichose. Material examined. Holotype: COSTA RICA: Alajuela: Rio San Lorencito, San Ramón Forest Reserve, 5 km N Colonia Palmareña, 900 m a.s.l., iii.1990, Curso Carabidae (♂, MNCR). Paratypes: COSTA RICA: Alajuela: same data as holotype (1 ♀, MNCR); Finca San Gabriel, 2 km SE Dos Rios, 600 m a.s.l., v.1990, II Curso Parataxon (1 ♀, MNCR); Puntarenas: Monteverde Biological Reserve, 1500 m a.s.l., cloud forest, 11–13.vi.2000, S.A. Marshall (2 ♂, 1 ♀, DEBU, MNCR); Monteverde Biological Station, 1500 m a.s.l., white pans in kitchen compost, 13.vi.2000, M. Buck (1 ♀, DEBU). PANAMA: Panamá Oeste: Cerro Campana, 5.vii.1974, C.W. & L. O’Brien & Marshall (1 ♂, CAS). Type locality. Costa Rica, Alajuela, Rio San Lorencito, San Ramón Forest Reserve. Etymology. The specific name (from the Latin ‘valgus’: bent out) refers to the distinctively bent surstylus.