Yoyetta Moulds 2012

Morphological key to species of Yoyetta cicadas Specimens must be set with wings spread to begin using this key.A vernier calliper is required to check measurements. Male specimens must often be examined microscopically and may require dissection in some instances. Key to males 1. Hind wing plagas white................................................................................ 2 - Hind wing plagas pale brownish, pinkish or grey........................................................... 19 2. Abdominal tergites 2–8 and sternites II–VII uniformly black or brown without orange-brown or yellow-brown markings; intersegmental membranes dark and inconspicuous.......................................................... 3 - Abdominal tergites 2–8 mainly black with contrasting orange-brown or yellow-brown markings, or contrasting intersegmental membranes; sternites II–VII mainly yellow-brown or orange................................................... 5 3. Fore wing basal membranes orange or pale orange-white............................................... Y. timothyi - Fore wing basal membranes red.......................................................................... 4 4. When viewed from ventral side, tip of aedeagus strongly bifurcate, with apical arms splayed laterally, sometimes in a weak “v” shape, at an angle of 150–180 degrees............................................................. Y. denisoni - When viewed from ventral side, tip of aedeagus undivided or weakly divided, without distinct lateral arms...... Y. kershawi 5. Tergites 5–7 partly black with contrasting, orange or yellow markings........................................... 6 - Tergites 5–7 entirely dark brown to black with yellow or orange coloration restricted to the intersegmental membranes.... 11 6. Opercula mainly pale grey to dark grey-brown.............................................................. 7 - Opercula mainly red or orange.......................................................................... 10 7. Dorsal surface of head, pronotum and mesonotum typically covered with dense gold pubescence; fore wing basal membranes orange or pink........................................................................................ 8 - Head, pronotum and mesonotum not covered in conspicuous pubescence; fore wing basal membranes pale grey.......... 9 8. Hind wing plagas narrow on ac2(v) (along vein 3); apex of theca blunt and club-like..................... Y. abdominalis - Hind wing plagas broad on ac2 (along vein 3), though not quite as broad as on jugum; apex of theca tapered and angled 30° ventrally.................................................................................... Y. loftyensis 9. Lateral sides of tergites 3–7 mainly orange; tergite 8 with orange markings on anterodorsal side................. Y. aaede - Lateral sides of tergites 3–7 mainly black (with yellow dorsolateral markings); anterodorsal side of tergite 8 black..................................................................................................... Y. serrata 10. Body length <23 mm; sternites III–VII bright reddish-orange without a continuous dark brown to black central marking............................................................................................... Y. spectabilis - Body length> 23 mm; sternites II–VII yellowish-orange or reddish-brown with a continuous dark brown to black central marking....................................................................................... Y. regalis 11. Fore wing length ≤ 26 mm ............................................................................. 12 - Fore wing length> 26 mm ............................................................................. 14 12. Opercula mainly black on basal half, pale dull yellow over remainder; meracantha orange..................... Y. incepta - Opercula almost entirely pale reddish-brown or pale grey; meracantha pale brown................................. 13 13. Pronotum dark brown to black with contrasting yellow-brown central marking; fore wing basal membranes orange.................................................................................................... Y. electrica - Pronotum dark brown to black without contrasting yellow-brown central marking; fore wing basal membranes grey to smoky orange.................................................................................... Y. hunterorum 14. Dorsal surface of head, pronotum and mesonotum with an inconspicuous, sparse covering of pubescence; lateral depressions adjacent to cruciform elevation conspicuous, brown to pale brown....................................... Y. grandis - Dorsal surface of head, pronotum and mesonotum typically with dense, black pubescence; lateral depressions adjacent to cruciform elevation inconspicuous, typically dark brown to black.............................................. 15 15. Hind wing plagas broad; with opaque white area expressed equally in cells ac3 and ac2(v) (either side of the jugal fold)............................................................................................... Y. subalpina - Hind wing plagas narrow or predominantly expressed in ac3, with a narrow extent on ac2(v) (along vein 3a)............ 16 16. Tergite 8 black with a brown, reddish-brown or dark reddish-brown posterior lateral marking on each side; apex of theca distinctly club-shaped; transparent flange along margin of recurvature not quite as broad as thecal shaft........... Y. regalis - Tergite 8 entirely dark brown to black without posterior lateral markings; apex of theca not club-shaped but narrow with ornamentation; transparent flange along margin of recurvature much broader than thecal shaft....................... 17 17. Hind wing plagas broad and occupying the jugum................................................... Y. douglasi - Hind wing plagas narrow with jugum mainly hyaline........................................................ 18 18. Fore wings conspicuously broad, with bend in ambient vein about equal to bend in costa; sternites III–VII typically with dark brown to black central markings................................................................. Y. ngarabal - Fore wings not conspicuously broad, with bend in ambient vein more pronounced than bend in costa; sternites III–VII usually without dark brown to black central markings........................................................ Y. verrens 19. Median lobe of uncus long, protruding conspicuously further ventrally than the claspers............................ 20 - Median lobe of uncus short, not protruding conspicuously further ventrally than the claspers......................... 23 20. Abdomen with sternite VIII entirely orange-brown or light orange brown............................. Y. darug sp. nov. - abdomen with sternite VIII dark brown to black at base grading to orange brown, becoming paler towards apex......... 21 21. Fore wings with weak to moderate infuscations along crossveins r and r-m; median lobe of uncus shorter than distance between dorsal beak and apex of upper pygofer lobe.................................................... Y. fumea sp. nov. - Fore wings with or without infuscations along crossveins r and r-m (a weak infuscation may be present); median lobe of uncus about as long as the distance between the apices of the upper pygofer lobe and dorsal beak.......................... 22 22. Median lobe of uncus narrow at base, widest around mid length and tapering towards apex................ Y. humphreyae - Median lobe of uncus narrow, slightly expanding distally, with rounded apex............................... Y. corindi 23. Head width <5 mm ................................................................................... 24 - Head width ≥ 5 mm ................................................................................... 33 24. Thorax mainly brown................................................................................. 25 - Thorax mainly black.................................................................................. 27 25. Tergite 1 pale brown; pronotum mainly pale brown...................................................... Y. celis - Tergite 1 black or dark brown; pronotum mainly brown or black............................................... 26 26. Fore wing length> 20 mm; cruciform elevation and surrounding areas adjacent to wing grooves all much paler and brighter than remainder of thorax.......................................................................... Y. tristrigata - Fore wing length <20 mm; cruciform elevation and surrounding areas adjacent to wing grooves not all distinctly paler than remainder of thorax........................................................................... Y. fluviatilis 27. Pronotum mainly brown, contrasting with mesonotum.................................. Y. tristrigata (dark specimen) - Pronotum mainly dark brown to black, not distinctly paler than mesonotum...................................... 28 28. Lateral edges of tergites orange, with dark markings along posterior margin (broad or narrow, often diffuse)............ 29 - Lateral edges of tergites orange, without dark markings on posterior margin...................................... 30 29. Fore wing length usually> 20 mm; theca recurved distally through 180 degrees; apex weakly upturned.......... Y. delicata - Fore wing length <20 mm; theca recurved distally through 120 degrees; apex downturned.................... Y. fluviatilis 30. Theca recurved distally through 180 degrees; apex straight or weakly upturned................................... 31 - Theca recurved distally through 120 degrees; apex straight or downturned....................................... 32 31. Fore wing length typically> 20 mm; apex of theca weakly upturned...................................... Y. delicata - Fore wing length <20 mm; apex of theca straight...........................................