Scarabaeus sakalava Montanaro & Tarasov 2024, sp. nov.

Scarabaeus sakalava Montanaro & Tarasov sp. nov. Materials Type status: Holotype. Occurrence: catalogNumber: http://id.luomus.fi/GZ.15827; recordedBy: I. Hanski group; individualCount: 1; sex: male; lifeStage: adult; occurrenceID: 16BD880A-0D65-5B0E-909A-A3DDD3A4D1A8; Taxon: taxonID: urn:lsid:biosci.ohio-state.edu:osuc_names:275502; scientificName: Scarabaeus sakalava; genus: Scarabaeus; specificEpithet: sakalava; scientificNameAuthorship: Montanaro & Tarasov; Location: country: Madagascar; stateProvince: Toliara; locality: Ambadira-Morondava; verbatimLocality: Ambadira-Mormdava; decimalLatitude: - 19.8299; decimalLongitude: 44.6403; georeferenceRemarks: coordinates inferred from locality name; Identification: identifiedBy: Giulio Montanaro; dateIdentified: 2023; Event: samplingProtocol: cattle dung; eventDate: 12-2006; Record Level: institutionID: http://grbio.org/cool/2vmj-fp0v; institutionCode: MZH; basisOfRecord: PreservedSpecimen Type status: Paratype. Occurrence: catalogNumber: http://id.luomus.fi/GZ.15828; recordedBy: I. Hanski group; individualCount: 1; sex: female; lifeStage: adult; occurrenceID: 54A6436B-20BF-5615-9B08-048D138D820E; Taxon: taxonID: urn:lsid:biosci.ohio-state.edu:osuc_names:275502; scientificName: Scarabaeus sakalava; genus: Scarabaeus; specificEpithet: sakalava; scientificNameAuthorship: Montanaro & Tarasov; Location: country: Madagascar; stateProvince: Toliara; locality: Ambadira-Morondava; verbatimLocality: Ambadira-Mormdava; decimalLatitude: - 19.8299; decimalLongitude: 44.6403; georeferenceRemarks: coordinates inferred from locality name; Identification: identifiedBy: Giulio Montanaro; dateIdentified: 2023; Event: samplingProtocol: cattle dung; eventDate: 12-2006; Record Level: institutionID: http://grbio.org/cool/2vmj-fp0v; institutionCode: MZH; basisOfRecord: PreservedSpecimen Type status: Paratype. Occurrence: catalogNumber: http://id.luomus.fi/GZ.15818; recordedBy: Ilkka Hanski; individualCount: 1; sex: male; lifeStage: adult; occurrenceID: 341C47FD-A05B-53A0-99AB-30725CE58997; Taxon: taxonID: urn:lsid:biosci.ohio-state.edu:osuc_names:275502; scientificName: Scarabaeus sakalava; genus: Scarabaeus; specificEpithet: sakalava; scientificNameAuthorship: Montanaro & Tarasov; Location: country: Madagascar; stateProvince: Toliara; locality: Marofandilia; decimalLatitude: - 20.1327; decimalLongitude: 44.5503; georeferenceRemarks: coordinates inferred from locality name; Identification: identifiedBy: Giulio Montanaro; dateIdentified: 2023; Event: samplingProtocol: fish baited trap; eventDate: 04-2004; habitat: dry deciduous forest corridor; Record Level: institutionID: http://grbio.org/cool/2vmj-fp0v; institutionCode: MZH; basisOfRecord: PreservedSpecimen Type status: Paratype. Occurrence: catalogNumber: http://id.luomus.fi/GZ.15817; recordedBy: Ilkka Hanski; individualCount: 1; sex: male; lifeStage: adult; occurrenceID: F6002F94-267E-5CBB-802A-F519E4BEC873; Taxon: taxonID: urn:lsid:biosci.ohio-state.edu:osuc_names:275502; scientificName: Scarabaeus sakalava; genus: Scarabaeus; specificEpithet: sakalava; scientificNameAuthorship: Montanaro & Tarasov; Location: country: Madagascar; stateProvince: Toliara; locality: Marofandilia; decimalLatitude: - 20.1327; decimalLongitude: 44.5503; georeferenceRemarks: coordinates inferred from locality name; Identification: identifiedBy: Giulio Montanaro; dateIdentified: 2023; Event: samplingProtocol: fish baited trap; eventDate: 04-2004; habitat: dry deciduous forest corridor; Record Level: institutionID: http://grbio.org/cool/2vmj-fp0v; institutionCode: MZH; basisOfRecord: PreservedSpecimen Type status: Paratype. Occurrence: catalogNumber: http://id.luomus.fi/GZ.15816; recordedBy: Ilkka Hanski; individualCount: 1; sex: male; lifeStage: adult; occurrenceID: 7722D749-3E83-50B3-BB60-38B86F7F2477; Taxon: taxonID: urn:lsid:biosci.ohio-state.edu:osuc_names:275502; scientificName: Scarabaeus sakalava; genus: Scarabaeus; specificEpithet: sakalava; scientificNameAuthorship: Montanaro & Tarasov; Location: country: Madagascar; stateProvince: Toliara; locality: Marofandilia; decimalLatitude: - 20.1327; decimalLongitude: 44.5503; georeferenceRemarks: coordinates inferred from locality name; Identification: identifiedBy: Giulio Montanaro; dateIdentified: 2023; Event: samplingProtocol: fish baited trap; eventDate: 04-2004; habitat: dry deciduous forest corridor; Record Level: institutionID: http://grbio.org/cool/2vmj-fp0v; institutionCode: MZH; basisOfRecord: PreservedSpecimen Type status: Paratype. Occurrence: catalogNumber: http://id.luomus.fi/GZ.57485; recordedBy: Sergei Tarasov; individualCount: 1; sex: female; lifeStage: adult; occurrenceID: 2E216A5E-330D-5C21-9711-BB32E3B096D8; Taxon: scientificName: Scarabaeus sakalava; genus: Scarabaeus; specificEpithet: sakalava; scientificNameAuthorship: Montanaro & Tarasov; Location: country: Madagascar; stateProvince: Toliara; locality: Morondava, Kirindy station; verbatimElevation: 71 m; verbatimCoordinates: - 20.066805, 44.657255; decimalLatitude: - 20.066805; decimalLongitude: 44.657255; georeferenceProtocol: GPS; Identification: identifiedBy: Giulio Montanaro; dateIdentified: 2023; Event: eventID: MG 22-29 a; samplingProtocol: human dung traps; eventDate: 03/ 07-03-2022; habitat: dry forest; Record Level: institutionID: http://grbio.org/cool/2vmj-fp0v; institutionCode: MZH; basisOfRecord: PreservedSpecimen Type status: Paratype. Occurrence: catalogNumber: http://id.luomus.fi/GZ.57486; recordedBy: Sergei Tarasov; individualCount: 1; sex: male; lifeStage: adult; occurrenceID: 055DDC5A-D731-5DB6-9A9C-B4359C2E84C5; Taxon: scientificName: Scarabaeus sakalava; genus: Scarabaeus; specificEpithet: sakalava; scientificNameAuthorship: Montanaro & Tarasov; Location: country: Madagascar; stateProvince: Toliara; locality: Morondava, Kirindy station; verbatimElevation: 71 m; verbatimCoordinates: - 20.066805, 44.657255; decimalLatitude: - 20.066805; decimalLongitude: 44.657255; georeferenceProtocol: GPS; Identification: identifiedBy: Giulio Montanaro; dateIdentified: 2023; Event: eventID: MG 22-29 a; samplingProtocol: human dung traps; eventDate: 03/ 07-03-2022; habitat: dry forest; Record Level: institutionID: http://grbio.org/cool/2vmj-fp0v; institutionCode: MZH; basisOfRecord: PreservedSpecimen Type status: Paratype. Occurrence: catalogNumber: http://id.luomus.fi/GZ.57487; recordedBy: Sergei Tarasov; individualCount: 1; sex: female; lifeStage: adult; occurrenceID: 64E02FAA-3180-51C1-9ACC-C9E42D0C3020; Taxon: scientificName: Scarabaeus sakalava; genus: Scarabaeus; specificEpithet: sakalava; scientificNameAuthorship: Montanaro & Tarasov; Location: country: Madagascar; stateProvince: Toliara; locality: Morondava, Kirindy station; verbatimElevation: 71 m; verbatimCoordinates: - 20.066805, 44.657255; decimalLatitude: - 20.066805; decimalLongitude: 44.657255; georeferenceProtocol: GPS; Identification: identifiedBy: Giulio Montanaro; dateIdentified: 2023; Event: eventID: MG 22-29 a; samplingProtocol: human dung traps; eventDate: 03/ 07-03-2022; habitat: dry forest; Record Level: institutionID: http://grbio.org/cool/2vmj-fp0v; institutionCode: MZH; basisOfRecord: PreservedSpecimen Type status: Paratype. Occurrence: catalogNumber: http://id.luomus.fi/GZ.57488; recordedBy: Sergei Tarasov; individualCount: 1; sex: female; lifeStage: adult; occurrenceID: 42053262-8E9F-5379-AD6C-6A6A57AA3507; Taxon: scientificName: Scarabaeus sakalava; genus: Scarabaeus; specificEpithet: sakalava; scientificNameAuthorship: Montanaro & Tarasov; Location: country: Madagascar; stateProvince: Toliara; locality: Morondava, Kirindy station; verbatimElevation: 71 m; verbatimCoordinates: - 20.066805, 44.657255; decimalLatitude: - 20.066805; decimalLongitude: 44.657255; georeferenceProtocol: GPS; Identification: identifiedBy: Giulio Montanaro; dateIdentified: 2023; Event: eventID: MG 22-29 a; samplingProtocol: human dung traps; eventDate: 03/ 07-03-2022; habitat: dry forest; Record Level: institutionID: http://grbio.org/cool/2vmj-fp0v; institutionCode: MZH; basisOfRecord: PreservedSpecimen Description Fig. 2 a, b male organism Catalog Number http://id.luomus.fi/GZ.15827 male organism has role in modeling TU denotes species species Parent Name Usage ID https://www.gbif.org/species/9074838 species Taxon ID http://zoobank.org/ 7AD8F87F-E7C1-4094-BD63-7662F167E9CB male organism: ovate; male organism, chitin-based cuticle chitin-based cuticle, lateral side encircles cuticular seta: red brown; chitin-based cuticle, ventral side encircles cuticular seta: red brown; chitin-based cuticle: black; chitin-based cuticle: glistening; male organism, antenna with 9 antennomeres antenna with 9 antennomeres, antennal club antennal club, flagellomere 5: present; antennal club, flagellomere 6: present; antennal club, flagellomere 7: present; antenna with 9 antennomeres: red brown; male organism, insect leg encircles cuticular seta: red brown; male organism, genal margin genal margin, posterior region: right angle to; genal margin, anterior region anterior region: protruding; anterior region: acute angle to; male organism, head margin at genoclypeal sulcus: notched; male organism, lateral clypeal tooth 1: acute angle to; male organism, lateral clypeal tooth 2: acute angle to; male organism, clypeal margin between clypeal teeth 1 clypeal margin between clypeal teeth 1: acute angle to; clypeal margin between clypeal teeth 1: notched; male organism, clypeal margin between clypeal teeth 1 and 2 clypeal margin between clypeal teeth 1 and 2: acute angle to; clypeal margin between clypeal teeth 1 and 2: notched; male organism, frons frons, cuticular carina cuticular carina: bilaterally paired; cuticular carina: transverse orientation; cuticular carina: decreased length; frons: protruding; male organism, vertex, granulated cuticle: present; male organism, gena gena, medial region, granulated cuticle: present; gena, lateral region: wrinkled; male organism, clypeus, anterior region: wrinkled;