Triepeolus obliteratus Graenicher 1911

Triepeolus obliteratus Graenicher, 1911 Figs 3B, 7B, 9B, 12G, 22 Triepeolus obliteratus Graenicher, 1911: 242 (♀, ♂), lectotype presently designated. Triepeolus obliteratus – Mitchell 1962: 477 (redescription). — Veit et al. 2021: 139 (new state record, new host association). Proposed common name Obliterated triepeolus. Diagnosis The following morphological features in combination tell T. obliteratus apart from all other Triepeolus in the simplex species group: the mesopleura laterally do not have erect/suberect, simple setae or have only sparse, short (<¼ MOD), erect/suberect, simple setae in addition to the usual appressed, branched setae (Figs 9B, 22A, C); the legs, at least from coxae to femora, are partially to predominantly dark brown/black (Figs 3B, 9B, 22A–C); the trochanters are normal (i.e., not tuberculate) (Fig. 9B); the dorsum of the mesosoma and metasoma have bands of pale-gray to white/off-white tomentum (Fig. 22); and the T1 discal patch is transversely oblong or elliptic (Fig. 22B). Triepeolus obliteratus most closely resembles T. oblongimacula sp. nov. and T. rhododontus, but in the latter two species the dorsum of the mesosoma and metasoma have bands of pale-yellow tomentum, the legs may be entirely reddish orange from trochanters to tarsi, and the pseudopygidial area of the female may be partially to predominantly reddish orange (as opposed to entirely dark brown/black; Fig. 22D), and in T. rhododontus the axillae commonly (as opposed to never) have reddish tips. Males of T. obliteratus also closely resemble those of T. eliseae Rightmyer, 2017 and T. michiganensis Mitchell, 1962, but in the latter two species, which are not in the simplex species group, the pygidial plate may have a well-defined basal transverse ridge, in T. eliseae the propodeal triangle is mostly covered in dense, appressed pale setae (as opposed to glabrous; Fig. 7B), and in T. michiganensis the clypeus has a well-defined glabrous midline (absent in T. obliteratus; Fig. 3B). Etymology The specific epithet was inspired by the fore wings each having the second submarginal crossvein incomplete in the lectotype and lectoallotype of this species, as indicated by Graenicher’s (1911) remark that “[b]oth of the specimens have the second transverse cubital nervure more or less obliterated in both wings”. Material examined Primary type material USA • ♀, lectotype; Wisconsin, Yellow River, Burnett County; 28–31 Jul. 1909; MPM ENT6434. Secondary type material USA • 1 ♂, lectoallotype; Wisconsin, Prescott, Pierce County; 13–19 Jul. 1910; MPM ENT6435. DNA barcoded material Available. BIN: BOLD:AEE4216. Specimens examined and sequenced: CANADA – Manitoba • 1 ♀; Spruce Woods Provincial Park; 49.6649° N, 99.2774° W; 30 Jul. 2019; T.M. Onuferko leg.; BOLD sample ID: CMNTO_003; TOPC. – Saskatchewan • 1 ♀; Avonlea Badlands; 50.0246° N, 104.9831° W; 11 Jul.–1 Aug. 2019; RSM Bee Team leg.; BOLD sample ID: CCDB-38770 E 10; RSKM RSKM _ ENT_E-225802 • 1 ♀; Douglas Provincial Park (Elbow); 51.0467° N, 106.4423° W; 22 Jul. 2019; T.M. Onuferko leg.; BOLD sample ID: CMNTO_001; CMNC • 1 ♀; Herbert; 50.4195° N, 107.2189° W; 15 Aug. 2013; C.S. Sheffield et al. leg.; BOLD sample ID: CCDB-32640 H07; RSKM RSKM_ENT_E-110579 • 1 ♀; near Avonlea; 50.0241° N, 104.9831° W; 30 Jun.–7 Jul. 2017; C.S. Sheffield and R. Oram leg.; BOLD sample ID: CCDB-38770 F01; RSKM RSKM_ENT_E-185267 • 1 ♂; same data as for preceding; 30 Jun.–7 Jul. 2017; C.S. Sheffield, R. Oram, J. Kalyniuk, and G. Wihlidal leg.; BOLD sample ID: CCDB-38770 E11; RSKM RSKM_ENT_E-185259 • 1 ♂; same data as for preceding; 30 Jun.–7 Jul. 2017; C.S. Sheffield, R. Oram, J. Kalyniuk, and G. Wihlidal leg.; BOLD sample ID: CCDB-38770 E12; RSKM RSKM_ENT_E-185261. USA – Iowa • 1 ♀; Butler County; 42.6260° N, 92.6896° W; 19 Jul. 2019; M. Vandever leg.; BOLD sample ID: CMNTO_057; CSUC. – Massachusetts • 1 ♀; Ashburnham, Worcester County; 42.7050° N, 71.9270° W; 6 Aug. 2015; M.F. Veit leg.; BOLD sample ID: CMNTO_249; TOPC. – New Hampshire • 1 ♂; Crotched Mountain Trail (Greenfield), Hillsborough County; 22 Jul. 2016; M.F. Veit leg.; BOLD sample ID: CMNTO_250; TOPC. Non-barcoded material CANADA – Alberta • 2 ♀♀; 8.5 km E of Clyde, Westlock County; 54.1661° N, 113.5003° W; 21 Jul. 2010; M. Buck leg.; RAM pmae00116017, pmae00116018 • 1 ♀; same data as for preceding; 21 Jul. 2010; M. Buck and Widen leg.; RAM pmae00116618 • 1 ♀; 15 km SW of Edson; 53.4917° N, 116.6044° W; 21 Jul. 2021; M. Buck leg.; RAM pmae00160149 • 1 ♀; Bridge Lake Natural Area, Westlock County; 54.1772° N, 113.4886° W; 27 Jul. 2011; M. Buck leg.; RAM pmae00120258 • 1 ♂; Opal Dunes, Thorhild County; 54.0075° N, 113.2681° W; 24 Jul. 2009; M. Buck and Palumbo leg.; RAM pmae00111591 • 1 ♀; Ukalta Dunes, Lamont County; 53.9747° N, 112.5072° W; 6 Aug. 2010; S. Widen leg.; RAM pmae00117232 • 1 ♀; Wainwright Municipal District, Ribstone Creek Heritage Rangeland Natural Area; 52.7269° N, 110.3667° W; 8 Jul. 2009; M. Buck leg.; RAM pmae00111947. – Manitoba • 2 ♀♀; 19.6 km WSW of Virden; 49.8014° N, 101.1961° W; 23 Jul. 2021; O. Abel leg.; CMNC • 1 ♂; 2.5 km E of Portage Sand Hills Wildlife Management Area on Road 54 N, Rural Municipality of Portage la Prairie; 49.7990° N, 98.2170° W; 5 Jul. 2019; E. Hanuschuk leg.; WRME WRME_EJH02370 • 1 ♀; 4.5 km SE of Portage La Prairie, Rural Municipality of Portage la Prairie; 49.9300° N, 98.2620° W; 16 Jul. 2018; E. Hanuschuk leg.; WRME JBWM0391741 • 1 ♀; 5 mi W of Carberry; 9 Aug. 1958; N.B. Chillcott leg.; CNC • 1 ♂; 6.5 km E of Portage Sand Hills Wildlife Management Area on Road 54 N, Rural Municipality of Portage la Prairie; 49.7980° N, 98.1650° W; 16 Aug. 2019; E. Hanuschuk leg.; WRME WRME_EJH02720 • 1 ♂; 7 km E of Marchand, Rural Municipality of Reynolds; 49.4440° N, 96.2700° W; 22 Jul. 2020; J. Gibbs leg.; WRME WRME0507584 • 1 ♀; Bald Head Hills (13 mi N of Glenboro); 9 Aug. 1958; R.B. Madge leg.; CNC • 1 ♂; Birds Hill Provincial Park; 50.0100° N, 96.9090° W; 2 Jul. 2020; J. Gibbs leg.; WRME WRME0507876 • 1 ♀; Birds Hill Provincial Park; 50.0115° N, 96.9065° W; 12 Aug. 2017; J. Gibbs and G.Y. Nozoe leg.; WRME JBWM0362976 • 1 ♀; Linden, Rural Municipality of Taché; 49.6790° N, 96.8890° W; 9 Aug. 2019; S. Shukla-Bergen leg.; WRME WRME0501597 • 3 ♀♀; Oak Hammock Marsh, Rural Municipality of Rockwood; 50.1750° N, 97.1550° W; 26 Aug. 2020; J. Gibbs leg.; WRME WRME0508600, WRME0508602, WRME0508637 • 1 ♀; Rd 13E (Oak Hammock Marsh Wildlife Management Area), Rural Municipality of Rockwood; 50.1820° N, 97.1610° W; 7 Aug. 2019; E. Hanuschuk leg.; WRME WRME_EJH01940 • 1 ♂; Shilo; 49.8124° N, 99.5131° W; 12 Jul. 2011; K. Wareham and S. Patterson leg.; WRME JBWM0385547 • 1 ♀; Spirit Hills; 49.6600° N, 99.2900° W; 25 Aug. 1978; T.D. Galloway leg.; WRME JBWM0296424 • 1 ♀; Spirit Sands (Spruce Woods Provincial Park); 49.6600° N, 99.2800° W; 2 Jul. 2018; J. Gibbs leg.; WRME JBWM0379545 • 2 ♂♂; same data as for preceding; 6 Jul. 2019; J. Gibbs leg.; WRME JBWM0422817, JBWM0422819 • 1 ♂; St. Adolphe, Rural Municipality of Ritchot; 49.6550° N, 97.1140° W; 12 Jul. 2019; S. Shukla-Bergen leg.; WRME JBWM0425506 • 1 ♀; Ste. Agathe; 29 Aug. 1969; Boyle leg.; LEM • 1 ♂; Transcona; 8 Aug. ??24; G.S. Brooks leg.; CNC 427182 • 1 ♂; Vita, Rural Municipality of Stuartburn; 49.1490° N, 96.6460° W; 11 Jul. 2019; S. Shukla-Bergen leg.; WRME JBWM0425564 • 1 ♀; Winnipeg; 21 Aug. 1935; R.R. Lejeune leg.; WRME JBWM0296432. – Saskatchewan • 1 ♀; 21 mi W of Moose Jaw; 28 Jul. 1972; C.D. Michener leg.; KUNHM M.G.R. Database No. 584 • 1 ♀; NCC’s Buffalo Pound property; 50.7293° N, 105.5985° W; 3 Aug. 2021; C.S. Sheffield and K. Palmier leg.; RSKM RSKM_ENT_E-125601 • 1 ♀; Nipawin; 18 Jul. 1952; W.P. Stephen leg.; WRME JBWM0295628. USA – Illinois • 1 ♀; Richardson Wildlife Foundation, Lee County; 9 Jul. 1996; J.K. Bouseman leg.; INHS 441217 / 362391 • 1 ♂; Sand Ridge State Forest (4 mi SW of Forest City), Mason County; 15 Jul. 1982; E. Miliczky leg.; INHS 7784. – Massachusetts • 1 ♂; 1.1 mi SE of Pepperell (Groton), Middlesex County; 28 Jul. 2008; M.F. Veit leg.; TOPC WHD00000335 • 1 ♂; 1.8 mi NE of Townsend, Middlesex County; 28 Jul. 2008; M.F. Veit leg.; TOPC WHD00000334 • 1 ♀; Ashburnham, Worcester County; 42.7050° N, 71.9270° W; 9 Aug. 2014; M.F. Veit leg.; TOPC • 1 ♀; Groton, Middlesex County; 42.6410° N, 71.5275° W; 15 Aug. 2009; M.F. Veit leg.; TOPC WHD00000332 • 1 ♂; same data as for preceding; 15 Aug. 2009; M.F. Veit leg.; TOPC WHD00000337. – Minnesota • 1 ♂; 6 mi SW of Detroit Lakes, Becker County; 30 Jul. 1981; J.R. Powers leg.; EMEC M.G.R. Database No. 495. – New Hampshire • 1 ♀; Albany, Carroll County; 43.9944° N, 71.3694° W; 25 Jul. 2010; M.F. Veit leg.; TOPC WHD00000333 • 1 ♀; Ashuelot (power line row), Cheshire County; 42.7930° N, 72.4210° W; 9 Aug. 2017; M.F. Veit leg.; BIML • 1 ♀; Pelham, Hillsborough County; 42.7027° N, 71.3047° W; 30 Jul. 2009; M.F. Veit leg.; TOPC WHD00000331 • 1 ♂; same data as for preceding; 30 Jul. 2009; M.F. Veit leg.; TOPC WHD00000336. – New Jersey • 1 ♂; USNM M.G.R. Database No. 497 • 1 ♂; USNM M.G.R. Database No. 497. – North Carolina • 1 ♀; Black Mountains; AMNH Ac. 33827 /M.G.R. Database No. 598 • 1 ♀, T. junctus paratype; Fayetteville; 2 Oct. 1941; H.E. Evans leg.; CUIC Cornell U No. 3927/M.G.R. Database No. 527 • 2 ♀♀; Moore County; 8 Oct. 1964; B.S. Heming leg.; DEBU debu01089133, debu01089136 • 1 ♀; same data as for preceding; 8 Oct. 1964; J.F. Cornell leg.; DEBU debu01089130 • 1 ♀; Swannanoa; 3 Sep. 1924; J.C. Crawford leg.; USNM M.G.R. Database No. 528. – North Dakota • 1 ♀; 13 mi NW of Inkster, Grand Forks County; 17 Jul. 1955; W.E. LaBerge leg.; KUNHM M.G.R. Database No. 585 • 1 ♀; 13 mi NW of Inkster, Grand Forks County; 17 Jul. 1956; W.E. LaBerge leg.; KUNHM M.G.R. Database No. 599 • 1 ♀; Grands Forks; 3 Aug. 1912; O.A. Stevens leg.; USNM 3404 /M.G.R. Database No. 586 • 1 ♀; Perth; 20 Aug. 1910; O.A. Stevens leg.; AMNH 6480 /M.G.R. Database No. 587. – Oklahoma • 1 ♀; Fort Cobb Wildlife Management