Dacimita David & Hancock 2022

Dacimita and Monacrostichus Dacimita and Monacrostichus share fuscous notopleural lobes, wing vein R 2+3 strongly bowed and running close to vein R 4+ 5 in basal portion and distinctly angled to run close to costa in apical portion, a short, broad, posteriorly rounded surstylus and similarly-shaped abdomen. The unusual shape of vein R 2+3 (synapomorphy) indicates a close relationship and it is likely that Monacrostichus evolved from Dacimita after dispersing into Southeast Asia, accompanied by a host plant shift to the newly available Citrus and characterised by its convoluted and sausage-like spermathecae, large and elongate pterostigma, strongly bowed vein M basal to r-m crossvein and elongate antennae with a bare arista. Dacimita has a swollen, rounded apex to its weakly coiled spermathecae (densely coiled and convoluted in Dacus and Bactrocera), a short and narrow pterostigma, weakly bowed vein M basal to r-m crossvein and a short-plumose arista. The gastrozonine genus Enicoptera Macquart also has a strongly bowed vein M basal to r-m crossvein but vein R 2+3 runs parallel to vein R 4+ 5 in basal portion, is distinctly arched subapically and strongly bowed away from costa in apical portion (Hancock and Drew 1999); it also has a semicircular scutellum and typically gastrozonine setation and is not related.