Helicosina crena Kuwahara & Marshall & Yau 2024, sp. nov.

Helicosina crena Marshall sp. nov. (Figs. 2A–B, 3A, 4, 7, 8) Description. Body length: Males 1.5–1.8 mm, females 1.6–1.8 mm. General (Fig. 2A): Male fore and hind femora slightly thickened (length 1.9–2.0 × and 4.4–5.6 × width, respectively). Male abdomen (Fig. 4): S5 broad, desclerotized posteromedially, and uniformly long-setose. Posterior opening of epandrium triangular (medial height subequal to ventral width); subanal plate long and very narrow. Surstylus folded into inner and outer lobes; outer lobe rounded and setose, inner lobe flat with a few marginal setae and a large, transparent, posteroventral scale. Cercus broad, lobate, narrowly fused dorsally. Distiphallus short-cylindrical with a very broad, bilobed opening subtended by broad, flap-like distal lobes. Material examined. Holotype: COSTA RICA: Puntarenas: Monteverde Biological Reserve, 1500 m, cloud forest, 11–13.vi.2000, S.A. Marshall (♂, MNCR). Paratypes: COSTA RICA: Alajuela: Rio Penas Blancas, 700 m, 18.viii.1986, L. Masner (1 ♂, DEBU). San José: Moravia, Zurquí de Moravia, 10˚02’58”N 84˚00’57”W, 1600 m, Malaise trap #1, 5–8.ix.2012, W. Porras (1 ♀, MNCR); Moravia, Zurquí de Moravia, 10˚02’58”N 84˚00’57”W, 1600 m, mixed methods, 5–6.viii.2013, S.A. Marshall (1 ♀, MNCR); Moravia, Zurquí de Moravia, Tower path, 10˚02’58”N 84˚00’57”W, 1600 m, Malaise trap #1, 24–31.v.2013, Proyecto ZADBI (1 ♂, MNCR); Moravia, Zurquí de Moravia, Tower path, 10˚02’58”N 84˚00’57”W, 1600 m, Malaise trap #1, 14–21.vi.2013, Proyecto ZADBI (1 ♂, MNCR); Moravia, Zurquí de Moravia, Tower path, 10˚02’58”N 84˚00’57”W, 1600 m, Malaise trap #1, 28.vi–5.vii.2013, Proyecto ZADBI (1 ♀, MNCR); Moravia, Zurquí de Moravia, Tower path, 10˚02’58”N 84˚00’57”W, 1600 m, Malaise trap #1, 12– 19.vii.2013, Proyecto ZADBI (2 ♀, MNCR); Moravia, Zurquí de Moravia, Tower path, 10˚02’58”N 84˚00’57”W, 1600 m, Malaise trap #1, 9–16.viii.2013, Proyecto ZADBI (1 ♂, MNCR); Moravia, Zurquí de Moravia, Tower path, 10˚02’58”N 84˚00’57”W, 1600 m, Malaise trap #1, 30.viii–6.ix.2013, Proyecto ZADBI (1 ♂, 1 ♀, MNCR); Moravia, Zurquí de Moravia, Tower path, stingless bee entrance, 10˚02’58”N 84˚00’57”W, 1600 m, mixed methods, 9.viii.2013, B. Brown (1 ♂, MNCR). Type locality. Costa Rica, Puntarenas, Monteverde Biological Reserve. Etymology. The specific name (from the Latin ‘crena’: incision, notch) is a noun in apposition and reflects the conspicuously notched male sternite 5.