Culladia Moore 1886

Culladia Moore, 1886 Culladia Moore, 1886: 382. Type species: Araxes admigratella Walker, 1863 = Araxes * Walker, 1863 b: 192. Type species: Araxes admigratella Walker, 1863 = Crambidion Mabille, 1900: 748. Type species: Crambidion achroellum Mabille, 1900. Viette 1990: 85 (syn.) = Nirmaladia Rose, 1983: 172, 175, figs. 1–7. Type species: Culladia dentilinaelis Hampson, 1919. M. Shaffer, Nielsen & Horak 1996: 183 (syn.) DIAGNOSIS The genus Culladia was revised by Bleszynski (1970a). Species of Culladia exhibit small, greyish wings. Vein R5 is very short or missing as observed in a few small Crambini genera (Bleszynski, 1970a; Léger et al., 2019). In male genitalia, the bifid sclerotized apex of the phallus is a potential apomorphy for the genus. Male genitalia exhibit the following characters: uncus and gnathos slender, with apex pointing downwards; valva slender, with costal arm at base, ventral process absent; pseudosaccus present; juxta short, with deep notch; vesica with one or more cornuti, with characteristic bifid sclerotized apex (Bleszynski, 1970a; pers. obs.). Female genitalia exhibit the following characters: papillae anales not coalesced dorsally and ventrally; posterior apophyses 1-2 X length of tergite VIII; anterior apophyses very short; antrum often with pair of sclerites; membranous pouch branching near antrum found in several species; corpus bursae usually with long, scobinate signum (Bleszynski, 1970a; pers. obs.) DISTRIBUTION Occuring in the Afrotropical, Oriental, and Australasian regions. Five species are reported from the Philippines: Culladia evae Bleszynski, 1970, C. hastiferalis (Walker, 1865), C. pseudoscoparia sp. n., C. suffusella Bleszynski, C. tonkinella Bleszynski, 1970. PHYLOGENETIC RELATIONSHIPS Culladia is sister to the Holarctic genus Pediasia Hübner, 1825 (Bleszynski, 1970a; Léger et al., 2019).