Eudonia Billberg 1820

Eudonia Billberg, 1820 Eudonia Billberg, 1820. Type species: Phalaena mercurella Linnaeus, 1758. = Boiea * Zetterstedt, 1839: 995. Type species: Phalaena mercurella Linnaeus sensu Zetterstedt, 1839. = Borea Stephens, 1852: 2 = Dasyscopa Meyrick, 1894: 464. Type species: Dasyscopa homogenes Meyrick, 1894. Léger et al., 2019, p. 769 (syn.) = Dipleurina Chapman, 1912: 507. Type species: Phalaena crataegella Linnaeus sensu Hübner, 1796. Nuss, 1999, p. 59 (syn.) = Dipluerina Sharp, 1913: 357 = Dipleurinodes P. Leraut, 1989: 14–16. Type species: Dipleurinodes mineti Leraut, 1989. Léger et al., 2019, p. 769 (syn.) = Eudipleurina P. Leraut, 1989: 10–11. Type species: Eudoria (Dipleurina) ankaratella Marion, [1957]. Léger et al., 2019, p. 769 (syn.) = Malageudonia P. Leraut, 1989: 20. Type species: Witlesia malgassicella Marion, 1956, by original designation. Nuss, 1999, pp. 21, 22–23, 59 (syn.) = Vietteina P. Leraut, 1989: 36. Type species: Vietteina ivelonensis Leraut, 1989. Nuss, 1999, p. 59 (syn.) = Witlesia Chapman, 1912: 507. Type species: Eudorea pallida Curtis, 1827. Munroe 1972: 47 (syn.) = Wittlesia Chapman, 1912: 507, 518, pl. 40, 44 DIAGNOSIS Descriptions of Eudonia are provided in Nuss (1999) and Li, Li & Nuss (2012). The presence of an appendix bursae in female genitalia is an apomorphy for Eudonia + Scoparia (Léger et al., 2019). The gnathos tip directed posterad or upwards as well as the absence of cornuti on vesica in male genitalia is apomorphic for Eudonia (Léger et al., 2019; Nuss, 1999). The following characters of male genitalia further characterize the genus: uncus broad and short; valva rather simple, lacking the free distal process of the sacculus. In female genitalia, the following characters are characteristic for Eudonia: anterior and posterior apophyses usually long (over 3 X tergite length); intersegmental membrane IX-X long (over 2 X tergite length), colliculum long, tubular, sclerotized; ductus bursae long, always membranous, corpus bursae with a signum, devoid of appendix bursae (Léger et al., 2019; W. Li et al., 2012). DISTRIBUTION Distributed on all continents and many oceanic islands, including subantarctic islands (W. Li et al., 2012; Nuss et al., 2003 –2023). Two species are found in the Philippines: Eudonia penicula sp. n. and Eudonia barbipennis (Hampson, 1897). PHYLOGENETIC RELATIONSHIPS Eudonia is sister to Scoparia (Léger et al., 2019).