Pseudoleiobunum Suzuki & Ohrui 1972

Pseudoleiobunum (superspecies maximum) japanense japonicum Roewer, 1957 Pseudoleiobunum japonicum Suzuki, 1940: 95. Leiobunum suzukii Roewer, 1957: 343, Figures 12 –13 in Tafel 26 (Type: 2♀ from Yokohama). Leiobunum bifrons Roewer, 1957: 343, Figure 14 in Tafel 26 (Type 1♀ from Nagasaki). Metaliobunum japonicum Suzuki, 1940: 276–279; 1960: 22; 1965: 353. Leiobunum japanense japonicum Suzuki, 1976: 202–204, Figures 17–22, 96, 147–160; Tsurusaki & Sasaji, 1991: 11, Figure 5B; Suzuki, 1986:26, Figures 30–31; Kato et al., 2023: 63. Leiobunum japanense Tsurusaki, 2006: 163, Figure 8B. Leiobunum japanense suzukii Crawford, 1992: 30. Materials examined. (NT. = N. Tsurusaki). JAPAN, MIE PREF.: Owase City: Yanokawa, Mt. Takamine, 400 m, 2 juv., 25 March 1989, Y. Nishikawa; NE of Yanokawa Pass, 310–380 m, 3 juv., 26 March 1989, Y. Nishikawa — JAPAN, SHIGA PREF.: Otsu City, Katsuragawa-Sakashita-cho, 450 m, 1 juv., 3 October 2011, N. Nunomura; Koka City, Tsuchiyama-cho, Left Bank of Nosugawa Dam, 410 m, 1♀, 27 May, 2012, NT.; Maibara City, Kozuwara, Oku-Ibuki Skiing ground [N35°31.078', E136°22.5867'], 635 m, 1♂, 9 June 2017, NT. — JAPAN, KYOTO PREF.: Fukuchiyama City, Yakuno-cho, Oku-Mizusawa, 1♂ 2♀, 13 July 2002, M. Sasakawa — JAPAN, OSAWA PREF.: Kishiwada City, Mt. Izumi-Katsuragi, 850 m, 2 juv., 28 March 1997, NT.; Ibaraki City, Shobo, Left Bank of R. Ai, 80 m, 10♂ 24♀, 2 June 1994, Y. Nishikawa; Mino-o City: Todoromi, 9 juv., 7 May 2010, H. Anaze et al.; Mino-o Park, Sasagare-no-komichi, ca. 370 m, 3 juv., 26 March 1977, Y. Nishikawa; Higashi-Osaka City, Yamate-cho, Hiraoka Shrine, 1♂ 1♀, 29 May 2005, H. Tanaka — JAPAN, HYOGO PREF.: Shiso City, Haga-cho, Hitsujigadakiguchi, 1♀, 10 June 1990, K. Yamamoto; Shiso City, Oya-cho, Yokoiki-oku, 1♀, 2 August 1990, K. Yamamoto; 1 juv., 3 November 1990, K. Yamamoto — JAPAN, NARA PREF: Gojo City, Nishi-Yoshino-cho, Tentsuji Pass to Mt. Norikura [N34°14.051' E135°44.4341'], 850 m, 1 juv., 19 October 2015, NT.; Yoshino-gun, Totsukawa-mura, Mt. Shaka-ga-dake, Fudo-Kiyadani [N34°6.3368' E135°52.3868'], 1090 m, 3 juv., 20 October 2015, NT.; Kamikitayamamura, Mt. Oda-ga-hara, 1 juv., 26 May 1973, H. Minato; Tanabe City: Mt. Jogamori, 1♂ 2 juv., 4 June 2018, I. Matoba; Ryujin-mura, Mt.Gomadan, 1♂, 15 July2018, I. Matoba;Hongu-cho, Hatenashi Mountains, HigashinokawaOku, 1100 m, 1♀, 26 July 1978, H. Minato; Arida-gun, Aridagawa-cho, Mt. Higashidani, 1♂, 24 June 2017, I. Matoba; Mt. Toishi, 1♂, 4 July 2019, I. Matoba; Mt. Shirama, 1♂, 16 May 2019, I. Matoba; Kaiso-gun, Kiminocho: Ta, 250 m, 4♀, 28 February 1994, K. Katsura; Machida, Seminar House, 270 m, 1♂ 1♀, 19 May 2012, R. Yuasa; Nishi-Muro-gun, Ohto-mura, Hyakken Gorge, 3 juv., 2 May 1999, H. Minato; Higashi-Muro-gun, Kozakawacho, Hirai, 1♀, 9 August 1980, H. Minato; Nachi-Katsu-ura-cho, Seigantoji Temple, 340 m, 1♀, 28 July 1983, NT.; Iwade City, Negoro, Genki-no-mori, [N34°17.7167', E135°19.8347'], 1♀, 23 May 2021, Y. Yamamoto — JAPAN, TOTTORI PREF.: Tottori City, Kokufu-cho, Amedaki Falls, 460 m, 1♀, 1 June 1989, NT.; M. Murakami, K. Shimokawa, 440 m, 1 juv., 15 November 2007, NT.; Tottori City, Kawahara-cho, Gobara, 1 juv., 29 October 2010, Yachiyo Enginering; Ochidani, 50 m, 1♀, 27 May 1988, NT.; Mitaki Gorge, 560 m, 1♂, 24 May 1992, M. Murakam; Kitamura, along Midori-shigen Forest Road [N35°23.4038', E134°5.216'], 440 m, 1 juv., 27 March 2017, NT.; Tottori City, Saji-cho, Sanno-daki, Falls, 430 m, 1♂ 2♀, 3 June 2001, NT.; T. Adachi & S. Okada; KItadani, 830 – 1200 m, 1♀, 1 June 2002, NT.; Tottori City, Shikano-cho, Mt. Jubo, Kochi to Azogoe, 370 m, 1♂, 14 June 1992, H. Kunimoto.; Tottori City, Shikano-cho, Mt. Jubo, Obata Forest Road, 230 m, 1 juv., 2 May 1992, NT.; 230 m, 1 juv., 18 May 1992, NT.; 230 m, 1♀, 16 June 1992, NT.; Iwami-gun, Iwami-cho, Nagatani, 200 m, 1♀, 18 May 2004, NT. et al.; Kugami, Higashihama, Kanro Shrine [N35°35.9478', E134°21.6338'], 10 m, 1 juv., 28 March 2017, NT.; Yazu-gun, Wakasa-cho, Mt. Hyonosen, Hibiki-no-mori, 850 m, 1♀, 3 August 2003, T. Okada; Yazu-gun, Chizu-cho, Ashizu-kei, Mitaki Dam [N35°16.54', E134°20.4293'], 820 m, 1♂, 3 June 2003, NT.; Ute Pass, 600 m, 1 juv., 9 August 2005, NT.; Mitaki Dam, 720–740 m, 1 juv., 26 November 1989, NT. — JAPAN, SHIMANE PREF.: Izumo City, Tachikue-kyo, 40 m, 1♀, 15 July 1988, NT. — JAPAN, OAKYAMA PREF.: Mimasaka City, Shiramizu, 225 m, 2♀, 22 June 2015, M. Yamada — JAPAN, HIROSHIMA PREF.: Shobara City, Takano-cho, Oka-Ouchi, 1♀, 22 June 2001, S. Okada; Saijo-cho, Misaka, 5 juv., 8 September 2005, S. Nakamura; Oi Pass, Oi Pass Shrine, 640 m, 1♂, 6 August 1990, NT.; Hiroshima City, Asa-kita-ku, Uga-kyo, 2♀, 30 May 1976, NT.; Kure City, Niko-kyo, 50m, 1♂, 17 July 1990, NT.; Hatsukaichi City, Mt. Gokurakuji, 1♀, 25 May 1975, NT.; Yamagata-gun, Aki-Ota-cho, Sandankyo Gorge, 1♂ 1♀, 3 July 1977, NT. & H. Komatsu — JAPAN, TOKUSHIMA PREF.: Tokushima City, Hachimancho, Bunka-no-mori, 1 juv., 25 November 1990, T. Tanabe; Myozai-gun, Kamiyama-cho, Mt. Kumosozan [N33°5 4.7095 ', E134°17.3679'], 1120–1140 m, 2♂, 1 August 1997, NT.; Oe-gun, Kamikawa-ho, Mt. Kotsuzan, 1080–1100 m, 1♂ 1♀ 1 juv., 18 May 1998, NT.; Miyoshi City, Ochiai Pass, 1320 m, 1 juv., 13 October 1994, T. Tanabe; HigashiIya, Sano, 1180m, 1♀, 13 July 1991, M. Kamata — JAPAN, KAGAWA PREF.: Zentsuji City, Oasa-cho, Mt. Oasa, ca. 450 m, 1 juv., 14 November 2006, S. Mitani; Kan-nonji City, Mt. Umpenji, 850 m, 1 juv., 16 May 2019, S. Mitani; Sanuki City, Okuboji Temple, 450 m, 1♀, 15 June 2007, NT. & M. Takeuchi; Takamatsu City, Shionoe-cho, Mt. Otaki, 1♂ 1♀, 27 July 2003, S. Mitani; Nakatado-gun, Kotonan-cho, Hirakawa, 1♀, 2 July 2001, S. Mitani; Kotoshira-cho, Mt. Kotohira, 1♂ 1♀, 15 September, 2002, S. Mitani; Chunan-cho, Mt. Onoseyama, 1♂, 14 July 2003, S. Mitani — JAPAN, EHIME PREF.: Matsuyama City, Mt. Takanawa, 1 juv., 5 May 1972, NT.; 1♂, 17 July 2010, R. Ogawa; 960 m, 1 juv., 3 May 1993, NT.; 920 m, 1 juv., 4 April 1994, NT.; Matsuyama City, Mt. Hikijiyama, 900 m, 1♂, 11 August 2001, NT.; Mt. Sugitate, under Sugitate Bridge, 1 juv., 5 May 1970, NT.; YuyamaAonami, 2♂, 3 June 1973, H. Tsutsumi; Shuso-gun, Tambara-cho, Kurase, Iwane Shrine, 110 m, 1 juv., 23 April 2000, NT.; To-on City, Mt. Saragamine, Kambayashi, 1♂, 21 July 1972, NT. et al.; Kambayashi, Kambayashi Route, 1 juv., 3 May 1972, NT.; Kami-Ukena-gun, Kuma-kogen-cho, Mt. Saragamine, 1 juv., 23 April 1972, NT.; 3 juv., 29 April 1972, NT., Y. Yaeshima, A. Wake; 3 juv., 14 May 1972, NT & H. Yamashita; 1♂, 4 June 1972, NT. et al.; 950 m, 1 juv., 5 November 1974, NT.; 1♀, 21 May 1986, T. Tanabe; Kuma-kogen-cho, Teppoishigawa, 1♀, 13 August 2000, H. Ishikawa; Bansho-dani, 1♂, 20 August 2000, H. Ishikawa; Kuma-kogen-cho, Mt. Ishizuchi, Tchichigoya to Mt. Iwaguro, 1600 m, 1♂, 5 August 1982, NT.; Mt. Omogo, 1360–1500 m, 3♂ 2♀, 3 August 1982, Y. Sugino; Mt. Okawamine, 1250 m, 1 juv., 5 November 1993, NT.; 1400m, 1 juv., 5 November 1993, NT.; Kita-gun, Uchiko-cho, Oda-Miyama, 800 m, 3 juv., 6 October 1992, E. Yamamoto; 2 juv., 4 May 1994, E. Yamamoto; 800 m, 2 juv., 25 May 1993, E. Yamamoto; Koyayama, ca. 1200–1300 m, 3♂ 4♀, 12 June 1988, E. Yamamoto; Koyayama, 1♀, 2 August 1995, E. Yamamoto; Koyayama, 1♀, 5 August 1995, E. Yamamoto; Namakusa-dani, 8 juv., 1 May 1994, E. Yamamoto; Nakakusa-dani, 2♀, 25 June 1994, E. Yamamoto; Namakusa-dani, 1♀, 20 July 1994, E. Yamamoto; Nakmakusa-dani, 3 juv., 18 October 1994, E. Yamamoto; 1♂ 1♀, 10 June 1994, E. Yamamoto; Namakusa-dani, 1220 m, 4♀, 1 May 1994, NT. & C. Tsurusaki; Natural Forest, 800–900 m, 1 juv., 7 June 1988, E. Yamamoto; Shishigoe, 980 m, 1♀, 23 August 1994, E. Yamamoto; Namakusadani, 3♂ 8♀, 7 August 1995, E. Yamamoto; Namakusa-dani, 3♂ 1♀, 10 August 1995, E. Yamamoto; Masa-goya, 800m, 2 juv., 2 May 1994, NT.; 3♂ 4♀, 5August 1991, E.Yamamoto; 1♂ 1♀, 11August 1995, E. Yamamoto; Kita-gun, Uchiko-cho, Oda: Miyanotani, 400 m, 1 juv., 2 January 1995, E. Yamamoto; 1 juv., 1 April 1992, E. Yamamoto; 400 m, 1♂ 4♀, 10 August 1992, E. Yamamoto; Kita-gun, Hijikawa-cho, Yamatosaka, 1♂ 2♀, 22 June 2004, K. Uyeno; Kita-Uwa-gun, Kihoku-cho, Ogura, Odaru-Medaru Falls, 255 m, 2♂, 28 May 2013, NT. — JAPAN, KOCHI PREF.: Nagaoka-gun, Otoyo-cho, S. Ugi, Yasaka Shrine, 1 juv., 6 April 1994, Y. Murakami; Agawa-gun, Ino-cho, Nanokawagoe, 1♂, 22 June 2011, H. Ishikawa — JAPAN, FUKUOKA PREF.: Tagawa-gun, Soeda-cho, Mt. Hikosan [N33°29.1069', E130°55.8407'], 800 m, 2♂ 1♀, 17 June 2022, NT.; 1100–1199.5 m, 1♀, 30 July 1982, NT.; Asakura City, Mt. Koshozan [N33°29.0169', E130°43.5336'], 850 m, 1♂, 17 June 2022, NT. — JAPAN, NAGASAKI PREF.:: Iki-gun, Ishida-cho, Kuki, 2♀, 3 June 1990, M. Yamashita; Kuki, 1♂, 2 May 1991, M. Yamashita; Kuki, 3♂ 2♀, 25 May 1991, M. Yamashita; Kukishoku, 3 juv., 4 April 1994, M. Yamashita; Kukishoku, 3♂ 11♀, 18 May 1991, M. Yamashita; Kukishoku, 1♀, 8 May 1992, M. Yamashita; Kukishoku, 3♀, 10 May 1992, M. Yamashita; Unzen City, Unzen, Mt. Yadake, Hobaru, 730 m, 1 juv., 29 November 1993, NT. & C. Tusrusaki; Goto City, Is. Wakamatsu, Mt. Ryukan, 120 m, 1 juv., 7 February 2008, K. Okumura; Is. Fukue, Mt. Outou, 80 m, 1♂ 2♀, 9 May 2009, K. Okumura; Is. Fukue, Mt. Chichigadake, 150 m, 2 juv., 14 February 2009, K. Okumura; Is. Fukue, Mt. Chichigadake, 150 m, 5 juv., 2 March 2009, K. Okumura; Matsuura-gun, Shin-Kamigoto-cho, Is. Nakadori, Hifumi-daki Falls, 1 juv., 8 February 2009, K. Okumura — JAPAN, KUMAMOTO PREF: Kikuchi City, Kikuchi-keikoku, 1♂, 29 July 1970, M. Yoshikura — JAPAN, MIYAZAKI PREF.: Mito-gun, Nango-mura, Sasanotoge Pass, 1200 m, 1 juv., 15 November 2012, K. Nishi. Kobayashi City, Suki, along Ayaminami Forest Road, 400 m, 1 juv., 25 July 2012, K. Nishi; 1 juv., 25 December 2012, K. Nishi; Higashi-usuki-gun, Shiiba-son, Mt. Eboshi, 1470 m, 1♀, 16 June 2013, K. Nishi — JAPAN, KAGOSHIMA PREF.: Is. Yakushima, Arakawa, Collision trap, 1200 m, 1 juv., 30 October 2007, T. Yamauchi. Diagnosis. (Fig. 2A) General morphology similar to type species P. japanense japanense but differs in the shape