Siphonaria zelandica Quoy & Gaimard 1833

Siphonaria zelandica Quoy & Gaimard, 1833 (Figs 26F–O, 27C–E) Siphonaria zelandica Quoy & Gaimard 1833: 344, pl. 25; figs 17–18 (type locality: Nouvelle-Zélande [New Zealand]).— Lamarck 1839: 205; Catlow & Reeve 1845: 100; H. Adams & A. Adams 1855 (in 1853–58): 271; 1863: 271; Iredale 1915: 478; Hanley 1858b: 153; Jenkins 1983: 4, pls 1a–h, 2a–h, 6a–c. fig. 3a–d; Paul 1984: 28; Wells 1984: 53; Phillips et al. 1984: 78, fig. (unnumbered); Villiers & Hodgson 1987: 135; Short & Potter 1987: 122, pl. 60, fig. 6; Ludbrook & Gowlett-Holmes 1989: 612, fig. 11.29 j, k; Jansen 1995: 90, fig. 376; Wilson 2002: 172, text-fig.; Wells et al. 2003: 289; White & Dayrat 2012: 69; Colgan & da Costa 2013: 74; Dayrat et al. 2014: 268, fig. 5 B, C. Siphonaria bifurcata — Hanley 1858b: 151; Angas 1867: 232; Tenison Woods 1878b: 100; Paetel 1873: 173; 1883: 117; 1889: 428; Whitelegge 1889: 117; Hedley 1917b: M95, 1918: M95; Iredale 1924: 276; Hubendick 1955: 4; Berry 1977: 209, fig. 19; Short & Potter 1987: 122 (not S. bifurcata Reeve, 1856). Siphonaria australis — Lamarck 1836: 562; Hanley 1858b: 151; Hochlowski & Faulkner 1984: 3838 (not S. australis Quoy & Gaimard, 1833). Siphonaria baconi Reeve 1856: pl. 6, species 30 (type locality: Swan River, New Holland [Perth, WA, Australia]).— Hanley 1858b: 151; Paetel 1883: 178; 1889: 428; Angas 1865: 189; Tenison Woods 1878b: 100; Pritchard & Gatliff 1903: 221; Verco 1907: 104; 1912 b: 204; Iredale 1924: 276; Cotton & Godfrey 1932: 154; Macpherson & Chapple 1951: 142; Hubendick 1955: 129; Macpherson & Gabriel 1962: 264, fig. 302; Hodgkin et al. 1966: 27, pl. 8, fig. 4; Galindo 1977: 416; Mapstone 1978: 85; Jenkins 1981: 12, pl. 1 b, c; Roberts & Wells 1981: 72, pl. 30; Trew 1983: 4; Jenkins 1983: 5; Paul 1984: 28; Short & Potter 1987: 122; White & Dayrat 2012: 61. Siphonaria zealandiae Paetel 1889: 430 (invalid; incorrect subsequent spelling of zelandica). Siphonaria zebra Reeve 1856: pl. 5, species 22 [not species 21] (type locality: Port Jackson [Sydney Harbour], Australia).— Hanley 1858b: 153; Paetel 1889: 430; Hedley 1918: M95; Iredale,1924: 276; Hubendick 1955: 4; Galindo 1977: 416; Jenkins 1983: 5, 12, pl. 1 e–h; Trew 1983: 7; White & Dayrat 2012: 69. Siphonaria (Ductosiphonaria) baconi — Hubendick 1945: 70; 1946: 37, pl. 6, fig. 10, 11. Siphonaria luzonica — Adcock 1893: 11; Shikama 1964: 6, pl. 2, fig. 7 (not S. luzonica Reeve, 1856). Planesiphon elegans Iredale 1940: 441, figs 3–4 (type locality: Keppel Bay, Queensland).— Marcus & Marcus 1960: 122; Jenkins 1983: 5, 12, pl. 1 d, pl. 2 g –h; Short & Potter 1987: 122; White & Dayrat 2012: 63. Siphonaria (Ductosiphonaria) bifurcata — Hubendick 1945: 22, 70; 1946: 37, pl. 2, figs 9–13 (not S. bifurcata Reeve, 1856). Siphonaria (Siphonaria) elegans — Hubendick 1945: 29, 72, fig. 44, 46, 47. Siphonaria (Siphonaria) australis — Hubendick 1945: 72; 1946: 49, pl. 3, fig. 28–31 (not S. australis Quoy & Gaimard, 1833). Ductosiphonaria bifurcata — McAlpine 1952: 42, fig. 3 (not S. bifurcata Reeve, 1856). Siphonaria elegans — McAlpine 1952: 42. Siphonaria (Pachysiphonaria) zealandica Hubendick 1955: 129–136, figs 5–8 (invalid; incorrect subsequent spelling of zelandica). Ductosiphonaria baconi — Cotton 1959: 411. Ellsiphon denticulata — Cotton 1959: 411 (not S. denticulata Quoy & Gaimard, 1833). Planesiphon bifurcata — Iredale & McMichael 1962: 82 (not S. bifurcata Reeve, 1856). Siphonaria bifaciata Galindo 1977: 416 (invalid; incorrect subsequent spelling, not S. bifurcata Reeve, 1856). Material examined. Type material. Lectotype of Siphonaria zelandica Quoy & Gaimard, 1833, from Nouvelle Zealandie [New Zealand (in error for Australia, see Jenkins, 1983: 5)] (MNHN IM 2000-5055, Fig. 26F). Two paralectotypes, same data as lectotype (MNHN IM 2000-5135). Lectotype of Siphonaria baconi Reeve, 1856 from Swan River [Perth, WA] (NHMUK 1979164.1, Fig. 26J). Two paralectotypes of Siphonaria baconi Reeve, 1856. Same data as lectotype (NHMUK 1979164.2 and 1979164.3). Lectotype of Siphonaria zebra Reeve, 1856 from Port Jackson [on type label; ‘ Philippine Islands’ in Conch. Icon. incorrect—see Taxonomic remarks below] [Sydney, Australia], (NHMUK 1979168 /1, Fig. 26H). Three paralectotypes of Siphonaria zebra Reeve, 1856. Same data as holotype (NHMUK 1979168/2-4). Syntype of Planesiphon elegans Iredale, 1940 from Keppel Bay, Qld; coll. H. Bernhard, Sept 1935 and 1936 (AM C.103710, Fig. 26L). Syntypes of P. elegans Iredale 1940. Expanded syntype data (AM C.108528, 6d, locations on label of North Keppel Island and Emu Park; AM C.126707, 5d, location on label of Emu Park; MV F13845, 1d, location on label of Emu Park). Other, non-type material. Australia, Qld: Keppel Bay, 23°25’S, 150°55’E (AM C.103710 d); Emu Park, 23°15’S, 150°49’E (AM C.126707 5d); North Keppel Is & Emu Park 2, 3°25’S, 150°55’E (AM C.108528 6d). Caloundra Shelly Beach, 26°48’S, 153°9’E (AM C.391924 d). NSW: Fingal Bay nr Port Stephens, 32°44.990’S, 152°10.481’E, NSW09-1 (AM C.585559 3p); Broken Head Terrigal, 33°26.796’S, 151°27.030’E, NSW08-1 (AM C.585380 10p, C.585052 p [M098]); Terrigal The Skillion, 33°27.008’S, 151°27.122’E, NSW08-2 (AM C.585591 4p, C.585057 p [M222]); Wy-ar-gine Point Balmoral, 33°49.159’S, 151°15.195’E, NSW06-5 (AM C.585445 11p, C.585047 p [M155], Fig. 26I, C.585048 p [M156], C.585049 p [M157]); Bombo Kiama, 34°39.232’S, 150°51.649’E, NSW03-1 (AM C.585557 3p); Murunna Point Camel Head, 36°22.720’S, 150°04.766’E, NSW02-1 (AM C.595960 2p). Vic: Bear Gully, 38°53.519’S, 145°59.029’E, V07-3 (AM C.585572 3p); Caves, Inverloch 38°39.777’S, 145°40’871’E, V07- 2 (AM C.585359 p); San Remo, headland, 38°31.913’S, 145°22.209’E, V07-4 (AM C.585582 34p); San Remo, 38°31.489’S, 145°21.858’E, V07-1 (AM C.585358 p); Frankston, 38°09.236’S, 145°06.457’E, V06-1 (AM C.585651 5p); West Head Flinders, 38°28.883’S, 145°01.727’E, V06-3 0 (AM C.585717 8p); Point Lonsdale (nr Queenscliff), 38°17.276’S, 144°36.977’E, V05-1 (AM C.585608, 4 p); Roadknight Point, 38°25.707’S, 144°11.102’E, V04-1 (AM C.585474 16p); Marengo Rocks Apollo Bay, 38°46.772’S, 143°39.997’E, V03-1 (AM C.585650 5p); Crofts Bay, 38°35.363’S, 142°50.633’E, V01-3 (AM C.585607 4p, C.585284 p [M177]); Armstrong Bay, 38°21.012’S, 142°21.633’E, V01-2 (AM C.585726 9p). SA: Cape Northumberland, 38°03.503’S, 140°40.378’E, SA15-1 (AM C.585351 p); Cape Northumberland Port Macdonnell, 38°03.308’S, 140°39.398’E, SA15-2 (AM C.585723 9p, C.585223 p [M200]); Cape Thomas, 37°04.461’S, 139°44.659’E, SA14-1(AM C.585766 7p,C.584893d [R3032], C.584935 p [SK038], C.585218 p [M199]); Fisheries Bay Lands End, 35°37.999’S, 138°06.921’E, SA13-2 (AM C.585713 8p, C.584900 p [SK509], C.585216 p [SK024]); Groper Bay nr West Cape, 35°14.108’S, 136°49.883’E, SA10- 1 (AM C.585601 4p); Pondalowie Bay, 35°13.989’S, 136°49.892’E, SA10-2 (AM C.585473 16p, C.585492 p); Port Neill, 34°07.102’S, 136°21.271’E, SA06-1 (AM C.585710 8p); Port Moonta, 34°03.273’S, 137°33.592’E, SA09-1 (AM C.585647 5p); Salmon Point, 33°38.547’S, 134°51.916’E, SA04-2 (AM C.585688 7p); Whyalla, 33°02.539’S, 137°35.511’E, SA07-1 (AM C.585675 6p); Rocky Point, 32°12.250’S, 133°14.861’E, SA02-4 (AM C.5856877p); Ceduna, 32°07.438’S, 133°40.260’E,SA03- 2 (AM C.585707 8p); Denial Bay Ceduna, 32°05.886’S, 133°34.817’E, SA03-1 (AM C.585762 10p); Port Le Hunte Point Sinclair, 32°05.554’S, 132°59.476’E, SA02- 2 (AM C.585706 8p); Wandrilla Beach, nr Cape Nuyts 32°01.894’S, 132°16.052’E, SA01-1 (AM C.585522, 20 p). WA: Cave Point, 35°06.965’S, 117°54.080’E, WA62- 1 (AM C.584768 5p, WAM S74186 5p); Waterbay Point, 35°05.540’S, 117°56.963’E, WA62-2 (AM C.584684 2p, WAM S74187 2p); Peaceful Bay, 35°02.989’S, 116°55.769’E, WA60-7 (AM C.584744 4p, WAM S74183 4p); 35°02.865’S, 116°55.722’E, WA60-8 (AM C.585579 8p); Wilson Head Ocean Beach, 35°02.250’S, 117°19.894’E, WA61-1 (AM C.584730 10p, WAM S74185 5p); Cape Riche 34°36.213’S, 118°45.401’E, WA62-5 (AM C.584714 5p, WAM S74188 5p); Bremer Bay Boat Harbour, 34°25.613’S, 119°23.818’E, WA63-2 (AM C.584773 5p, WAM S74189 5p); Sarge Bay Cape Leeuwin, 34°22.091’S, 115°08.820’E, WA60-4 (AM C.585371 p); Augusta, 34°20.451’S, 115°10.069’E, WA60-5 (AM C.584699 3p, WAM S74182 3p); Alexander Bay 2, 33°53.467’S, 122°44.995’E, WA64-4 (AM C.584776 5p, 74191 5p); Alexander Bay, 33°53.374’S, 122°44.922’E, WA64-3 (AM C.584774 5p, 74190 5p); Salmon Beach Esperance, 33°53.254’S, 121°50.381’E, WA64-2 (AM C.585580 3p); Blue Haven Beach Esperance, 33°53.105’S, 121°51.687’E, WA64-1 (AM C.585314 p [M097]); Cowaramup Point, 33°51.934’S, 114°58.904’E, WA60-3 (AM C.585521 2p); Yallingup, 33°38.358’S, 115°01.481’E, WA60-9 (AM C.584696 3p, WAM S74184 2p); Point Dalling Dunsborough, 33°35.955’S, 115°06.315’E, WA59-4 (AM C.584750 5p, C.595950 p [M129], WAM S74180 5p); Point Casuarina Bunbury, 33°18.544’S, 115°38.201’E, WA59- 3 (AM C.585578 3p); groyne nr Robert Point Mandurah, 32°31.270’S, 115°42.409’E, WA59-1 (AM C.585437 5p, WAM S74179 5p); Fremantle Hbr breakwater 32°03.342’S, 115°43.987’E, WA58-5 (AM C.585612 4p); Longreach Bay point, Rottnest Is, 31°59.333’S, 115°32.063’E RI01 (AM C.584783 4p, WAM S74164 4p); Point Brown Swan River, 32°02.344’S, 115°45.471’E, WA59-5 (AM C.584709 4p, C.584899 p [SK039], C.585935 p [M127, SK243], WAM S74181 4p); Grey, 30°39.968’S, 115°08.072’E, WA58-2 (AM C.585367 p); Jurien Bay 30°17.244’S, 115°02.482’E, WA58-1 (AM C.585576 3p); Cambawarra Head Green Head, 30°04.136’S, 114°57.830’E, WA57-6 (AM C.585518 2p); Illawong, 29°42.254’S, 114°57.542’E, WA57-5 (AM C.584708 4p, WAM S74178 4p); Illawong bch rocks, 29°42.198’S, 114°57.551’E, WA57-4 (AM C.584705 4p, WAM S74177 3p); S end Leander Point Port Denison, 29°16.725’S, 114°54.918’E, WA57-2 (AM C.584682 2p, WAM S74176 2p); Leander Point Port Denison, 29°16.568’S, 114°54.858’E, WA57-1 (AM C.584692 3p, WAM S74175 2p); Cape Burney Geraldton, 28°52.084’S, 114°38.056’E, WA54-1 (AM C.584748 3p, WAM S74174 3p); Turtle Bay East Wallabi Is, 28°25.804’S, 113°44.538’E, WA55-1 (AM C.585658 5p); Horrocks, 28°21