Euura clitellata ZMUO.

141 Euura clitellata (Serville, 1823) Figs 138–141, 375–378 Nematus clitellatus Serville, 1823: 64. Holotype ♀ (MRSN), not examined. Type locality: France; vicinity of Paris. Best match: possibly ZMUO.029709 and ZMUO.040755, but wings slightly smoky. Nematus kirbyi Dahlbom, 1835a: 40, pl. 1 fig.4. Lectotype ♂ (ZMHB), designated by Haris (1997), not examined. One originally unlabelled male specimen (MZLU2017282) under the name “ Nematus kirbyi ” in Dahlbom’s collection (MZLU) could be a paralectotype. Type locality: Germany or Scandinavia. Synonymy by Taeger & Blank (1998). Nematus flaviventris Hartig, 1840: 24. Lectotype designated below. Synonymy by Taeger & Blank (1998). Nematus leucocarpus Hartig, 1840: 27. Lectotype designated by Taeger & Blank (1998). Synonymy by Taeger & Blank (1998). Nematus brachycercus Hartig, 1840: 25. Lectotype designated by Haris (1997). Syn. nov. Nematus squalidus Eversmann, 1847: 21. Lectotype designated below. Syn. nov. Nematus cinereus Eversmann, 1847: 18. Lectotype designated below. Syn. nov. Nematus contractus Eversmann, 1847: 20. Lectotype designated below. Synonymy with Pachynematus flaviventris by Konow (1905b). Nematus diaphanus Eversmann, 1847: 19–20. Lectotype designated below. Synonymy with flaviventris by Konow (1904d). Nematus fruticum Eversmann, 1847: 18. Lectotype designated below. Syn. nov. Nematus umbripennis Eversmann, 1847: 20–21. Lectotype designated below. Synonymy with kirbyi by Benson (1958a). Nematus griseus Eversmann, 1847: 18–19. Syntypes ♀♀ (DEI-GISHym30040–47, ZIN). Type locality: Russia, Kasan. Tentatively synonymised by Enslin (1916a). Like Nematus kirbyi Dahlbom. Tenthredo (Selandria) umbrosa Eversmann, 1847: 32. Lectotype designated below. Syn. nov. Epitactus praecox Förster, 1854b: 435–436. Lectotype designated by Haris (1997). Synonymy with kirbyi by Muche (1974). Nematus trisignatus Förster, 1854a: 292–294. Lectotype designated by Haris (1997). Synonymy by Enslin (1916a). Nematus microps Förster, 1854b: 421–422. Lectotype designated by Haris (1997). Synonymy by Haris (1997). Nematus pectoralis Snellen van Vollenhoven, 1867: 223–224. Lectotype ♀ (ZMAN), designated by Thomas (1987), not examined. Type locality: Netherlands, Brummen. Synonymy with trisignatus by Konow (1904c). Primary homonym of Nematus pectoralis Serville, 1823. Nematus turgidus Zaddach, 1876: 82–83. Syntypes ♀♀, probably destroyed (Blank & Taeger 1998). Type locality: Russia, Insterburg [Chernyakhovsk], and Poland, Gdansk [Danzig]. Synonymy by Enslin (1916a). Nematus tener Zaddach, 1884 in Brischke 1884: 128. Syntypes ♂♂, probably destroyed (Blank & Taeger 1998). Type locality: Poland, Gdansk [Danzig]. Synonymy by Konow (1904d). Pachynematus infumatus Marlatt, 1896a: 93 (key), 107–108. Synonymy with Pachynematus kirbyi (Dahlbom) by Smith (1979). Pachynematus zaddachi Konow, 1901c: 110. Lectotype designated by Haris (1997). Synonymy with umbripennis by Enslin (1916a). Pachynematus glesipennis Konow, 1903a: 377 (key). Lectotype designated by Haris (1997). Synonymy with umbripennis by Enslin (1916a). Pachynematus foveolatus Konow, 1903a: 379 (key). Lectotype ♂ (GBIF-GISHym4670, SDEI), designated by Haris (1997). Type locality: Russia, Irkutsk. Synonymy by Haris (1997). Pachynematus lentus Konow, 1903a: 382 (key). Lectotype designated by Haris (1997). Synonymy by Muche (1974). Pachynematus clitellatus var. transigens Enslin, 1916a: 479. Lectotype ♀ (DEI-GISHym84486, ZSM), designated by Haris (1997). Type locality: Germany, Neuss. Synonymy by Haris (1997). Pachynematus laevigatus var. flavissimus Enslin, 1916a: 483. Syntype (?) ♀ (DEI-GISHym84487, ZSM) [seems to match the original description]. Type locality: Germany, Oranienbaum. Synonymy by Taeger & Blank (1998). Pachynematus refractarius MacGillivray, 1921b: 31–32. Holotype ♀ (INHS), not examined. Type locality: USA, Maine, Orono. Synonymy with Pachynematus kirbyi (Dahlbom) by Smith (1979). Based on the original description it could be similar to Nematus kirbyi Dahlbom. Pachynematus truncatus Benson, 1948a: 60–65. Holotype ♂ (B.M.TYPEHYM.1.702, BMNH). Type locality: United Kingdom, England, Bucks., Slapton. Synonymy by Haris (1997). Secondary homonym of Pontania truncata Marlatt, 1896a [Euura truncata (Marlatt, 1896)]. Pachynematus smithae Ross, 1945: 106 (key), 116. Holotype ♂ (INHS), not examined. Type locality: USA, New Hampshire, Mt Washington. Syn. nov. Nematus (Pachynematus) omega Benson, 1955: 103–104. Lectotype designated below Syn. nov. Pachynematus aequalis Lindqvist, 1974a: 120. Syn. nov. Diagnosis Based on nuclear data, E. clitellata seems to be a complex of 3–5 species. Although we have not studied the holotype of clitellatus, based on the description given by Taeger & Blank (1998: 264) it could fit well, for example, with the specimen ZMUO.040755 that is extensively yellow, including hind tarsi. Specimens with that kind of colouration seem to be restricted to the nuclear cluster containing ZMUO.040755, which we have called E. clitellata sensu stricto (s. str.) (http://www.boldsystems.org/pics/LEFIJ/ZMUO.040755%2B1599793934.JPG). Generally, it seems that E. clitellata s. str. and the nuclear cluster 3 have yellow or paler hind tarsi compared to nuclear clusters 4 and 5, which have black or darker hind tarsi. Otherwise there seems to be little difference between the clusters and all of them vary widely in colouration (from mostly black to extensively pale) and habitus. There is also a nuclear cluster 2, represented by a single larva (ZMUO.040880). It is perhaps possible that E. clitellata s. str. and cluster 3 are conspecific, like the clusters 4 + 5, but it is very unlikely based on nuclear genes that E. clitellata s. str. + cluster 3 on one hand is conspecific with the cluster 4 + 5 on the other hand. In Europe, the clusters 3 and 5 seem to be restricted to northern Fennoscandia. For the cluster 4 (including perhaps cluster 5), the earliest available name might be Nematus brachycercus Hartig, 1840. Females of E. clitellata are very similar to E. fallax. If the mesoscutellum is partly pale, then there nevertheless seem still to be small differences in the colour pattern: – mesoscutellum with black midline along its length or pale dots laterally, if almost completely pale then with black dots anteriorly and posteriorly ..................................................................... E. fallax – mesoscutellum completely pale or anteriorly (partly) pale and posteriorly black ........... E. clitellata. Type material examined Lectotype Nematus flaviventris, here designated GERMANY • ♀; northern Germany [according to Hartig 1840: 22]; ZSM, DEI-GISHym3277. Notes Haris (1997) may have intended to designate this specimen as the lectotype, but according to Stephan M. Blank none of four syntypes was labelled as lectotype and the other labels as described by Haris in the lectotype designation do not exactly fit any of the specimens. Unfortunately, the specimen may be lost. It was not found in the ZSM, but photos of lateral and dorsal views are available. Not studied by Taeger & Blank (1998). Lectotype Nematus leucocarpus GERMANY • ♀; northern Germany [according to Hartig 1840: 22]; ZSM, DEI-GISHym84485. Notes 8.5 mm. The mesepisternum is completely pale, unlike any other specimen we have seen. Lectotype Nematus brachycercus GERMANY • ♀; northern Germany [according to Hartig 1840: 22]; ZSM, DEI-GISHym3208. Lectotype Nematus squalidus, here designated RUSSIA – Tatarstan • ♀; Kasan Province; 55.8° N, 49.1° E; E. Eversmann leg.; ZIN, DEI-GISHym30060. Notes Best morphological match: e.g., ZMUO.032426, ZMUO.039092, ZMUO.046267, ZMUO.045459, ZMUO.063011, DEI-GISHym31252, DEI-GISHym84266. Lectotype Nematus cinereus, here designated RUSSIA – Tatarstan • ♂; Kasan Province; 55.8° N, 49.1° E; E. Eversmann leg.; ZIN, DEI-GISHym30036. Notes Resembles Nematus kirbyi. Lectotype Nematus contractus, here designated RUSSIA – Tatarstan • ♀; Kasan / Ulyanovsk [Simbirsk] / Orenburg; ZIN, DEI-GISHym30052. Notes Similar to Nematus kirbyi. Paralectotype Nematus contractus RUSSIA – Tatarstan • ♂; Kasan / Ulyanovsk [Simbirsk] / Orenburg; ZIN, DEI-GISHym30053. Notes Perhaps Nematinus bilineatus (Klug, 1819). Lectotype Nematus diaphanus, here designated RUSSIA • ♀; Orenburg oblast; 51.8° N, 55.1° E; E. Eversmann leg.; ZIN, DEI-GISHym30051. Notes Similar to Nematus kirbyi. Lectotype Nematus fruticum, here designated RUSSIA – Tatarstan • ♂; Kasan Province; 55.8° N, 49.1° E; E. Eversmann leg.; ZIN, DEI-GISHym30035. Notes Similar to Nematus kirbyi. Lectotype Nematus umbripennis, here designated RUSSIA – Tatarstan • ♀; Kasan Province; 55.8° N, 49.1° E; E. Eversmann leg.; ZIN, DEI-GISHym30054. Notes Similar to Nematus kirbyi. Lectotype Tenthredo (Selandria) umbrosa, here designated RUSSIA – Tatarstan • ♂; Kasan Province; 55.8° N, 49.1° E; E. Eversmann leg.; ZIN, DEI-GISHym30048. Notes Similar to Nematus kirbyi. Lectotype Epitactus praecox GERMANY – Nordrhein-Westfalen • ♀; Aachen area; 50.77° N, 6.09° E; ZSM. Notes Similar to Nematus kirbyi. Lectotype Nematus trisignatus GERMANY – Nordrhein-Westfalen • ♀; Aachen area; 50.77° N, 6.09° E; ZSM. Lectotype Nematus microps GERMANY – Nordrhein-Westfalen • ♂; Aachen area; 50.77° N, 6.09° E; ZSM, DEI-GISHym12485. Holotype Pachynematus infumatus USA – Michigan • ♂; Ingham, Agricultural College; 42.746°N, 84.483°W; USNM, USNMENT00778520, http://n 2t.net/ark:/65665/322992bb4-e435-4a95-83cc-6b7a4796c66a. Notes Similar to Nematus kirbyi. Lectotype Pachynematus zaddachi GERMANY – Mecklenburg-Vorpommern • ♂; Kalkhorst; 53.32° N, 13.07° E; 1 Aug. 1889; F.W. Konow leg.; SDEI, GBIF-GISHym3875. Notes Similar to Nematus kirbyi. Lectotype Pachynematus glesipennis RUSSIA – Irkutskaja oblast • ♂; Irkutsk; 52.29° N, 104.24° E; SDEI, GBIF-GISHym3876. Lectotype Pachynematus foveolatus RUSSIA – Irkutskaja oblast • ♂; Irkutsk; 52.29° N, 104.24° E; SDEI, GBIF-GISHym4670. Lectotype Pachynematus lentus GERMANY • ♀; Thüringen; SDEI, GBIF-GIS