Tenomerga anguliscutis

Tenomerga anguliscutis (Kolbe, 1886) [Japanese name: Minami-nagahiratamushi (Kourai-nagahiratamushi)] Figs 1C, D, 2B, 3C, D, 4B, 8, 13B; Table 1 Cupes anguliscutis Kolbe, 1886: 200, pl. 11, fig. 33. Tamanuki 1928: 251; Atkins, 1963: 147. Tenomerga anguliscutis: Neboiss 1984: 451; Yoshitomi 2016: 183; Wang & Hájek 2018: 532. Cupes formosanus Tamanuki, 1928: 251. Atkins 1963: 147 (synonymized by Neboiss 1984). Type specimen examined. Holotype of Cupes formosanus Tamanuki, 1928 (Fig. 2B): male (SEHU), “ 24/VII 1925 Formosa Kono, Uchida”, “ Cupes formosanus Tamanuki Type”, “ Tenomerga anguliscutis (Kolbe) det. Neboiss 1983”, “0000001301 Sys.Ent Hokkaido Univ. Japan [SEHU]”, “HOLOTYPE Appended label by ÔHARA, INARI, KANBE SUZUKI and HIRONAGA 2007”. Other specimens examined. [JAPAN] <Tokara Isls.> 1 female, (EUMJ), Nakanoshima, 16. VII. 1961, Y. Hama leg.; 1 female, (EUMJ), Nakanoshima, 26. VI. 1962, H. Yokoyama leg. <Ryukyu: Okinawa-jima> 1 male, (EUMJ), Iji, Kunigami-son, 19. VI. 2008, M. Matsumura leg.; 1 male, (EUMJ), Isakawa-yama, 14. V. 1931; 1 male, (EUMJ), Kakazu, Tomishiro, 4. VI. 2015, Y. Tamaki leg.; 1 female, (KAC), Kenken, Motobu-cho, 20. VI. 2000, S. Inada leg.; 1 male, (EUMJ), Kenmin-no-mori, Onna-son, 24. VI. 2012, T. Kohama leg.; 1 male, (EUMJ), Kenmin-no-mori, Onna-son, 7. X. 2012, T. Kohama leg.; 1 female, (EUMJ), Kenmin-no-mori, Onna-son, 9. VI. 2012, T. Kohama leg.; 1 male, (EUMJ), Nago-dake, Nago-shi, 4. VII. 2017, J. Miyagi leg.; 1 male, (EUMJ), Nakadomari, Onna-son, 23. VI. 2016, T. Kuwayama leg. LT; 1 female, (KAC), Sedaka, Nago-shi, 17–21. V. 2011, T. Yamazaki leg.; 1 female, (EUMJ), Tahara, Itoman-shi, 6. V. 2019, H. Yoshitake leg. LT; 1 female, (EUMJ), Yakata-rindo, Onna-son, 30–31. VII. 2011, M. Osada leg. LT; 2 males, (EUMJ), Yamada-jyoshi, Onna-son, 15. VI. 2011, T. Kohama leg.; 4 males, (EUMJ), Yamada-jyoshi, Onna-son, 15–16. VI. 2011, M. Osada leg.; 1 male, (EUMJ), Yona, 26. VI. 1971, M. Kinjo leg.; 1 female, (SEHU), Yonaha-dake, 26. VI. 1973, T. Hatayama leg. (Nakane coll. 1999). <Ryukyu: Ishigaki-jima> 1 female, (EUMJ), Omoto-dake, 8. VII. 1964, Y. Hama leg. <Ryukyu: Iriomote-jima> 1 female, (KAC), Nakamagawa-rindo (20–50m), 18–23. V. 2010, K. Akita leg. FIT with LT; 1 female, (KAC), Otomi-rindo, 9. V. 1995, H. Onoda leg. [KOREA] 1 male, (NSMT), Seoul, 13. VI. 1936, E. Mizushima leg.; 1 male, (SEHU), Seoul, 21. VII. 1923, P. Gallois leg.; 1 female, (EUMJ), Seoul, 30. VI. 1933; 1 female, (SEHU), Seoul, VII. 1920; 1 female, (NSMT), Seoul, VII. 1963; 1 female, (SEHU), Cheuulfo, couif, 1924, mr. Glbois leg.; 1 female, (SEHU), Incheon Metropolitan City, 18. VII. 1923, Yuuki leg.; 1 male, (SEHU), no data, no data. Coll. R. Kano. [TAIWAN] 1 male (TARI), Holisha, 20. V. 1919. [CHINA] 3 males, (EUMJ), Shunyi Distr., Beijiang, III–IV. 2022; 1 male, (EUMJ), Guangzhou, 4. V. 2022, Hwang leg.; 1 male, (NSMT), Chinh sen, 11. VI. 1935, T. Tsuchiyama leg.; 1 male, (NSMT), Harbin, VIII. 1941, M. i. Nikitin leg.; 1 female, (SEHU), Dalian, 3. VII. 1929, N. Tomari leg.; 1 male, (NSMT), Dalian, 31. VII. 1937, N. Tomari leg. Diagnosis. Color of body yellowish brown, with alternating dark and pale brown markings. Elytra (Fig. 4B) with nine longitudinal rows of window punctures; window punctures round to square. Male (Fig. 1C). Eyes (Fig. 3C) large, prominent. Antennae long and slender, reaching about proximal 3/4 of elytra; AL/TL 0.75–0.88 (0.82); approximate ratio of each antennomere (n = 10; average) as 3.59: 1.00: 3.61: 4.28: 4.97: 5.25: 5.50: 5.47: 5.59: 5.45: 7.15. Anterior pronotal angles (Fig. 3C) acute or right-angled. HL/HW 0.53– 0.63 (0.58); PW/PL 1.27–1.44 (1.36); EL/EW 2.61–3.00 (2.81); EL/PL 5.38–6.36 (5.81); EW/PW 1.44–1.60 (1.52); TL/EW 3.54–3.94 (3.72); EYL/EYW 1.50–2.00 (1.69); TEL/HL 0.08–0.18 (0.14). Apex of sternite IX acuminate. Bifurcate process (Fig. 8D) with short median strut (Y-shaped in Hörnschemeyer, 2009). Parameres (Fig. 8A, B) with posterior projection of dorsal margin (pp) triangular; apical hook (Fig. 8C) bi-pointed; ventro-marginal spines (vms) long, reaching apical hook. Apical margin of dorsal plate (dp, Fig. 8A) deeply notched. Apical margin of mesal lobe (ml, Fig. 8A) excised. Female (Fig. 1D). Eyes (Fig. 3D) small. Antennae short and relatively stout, reaching about proximal 1/2 of elytra; AL/TL 0.51–0.70 (0.64); approximate ratio of each antennomere (n = 10; average) as 3.47: 1.00: 3.39: 3.68: 3.93: 4.05: 4.19: 4.13: 4.17: 4.07: 5.58. Anterior pronotal angles (Fig. 3D) acute or right-angled. HL/HW 0.60–0.68 (0.64); PW/PL 1.32–1.58 (1.41); EL/EW 2.55–3.51 (2.80); EL/PL 5.64–7.22 (6.10); EW/PW 1.47–1.68 (1.55); TL/EW 3.34–4.43 (3.66); EYL/EYW 1.67–2.33 (1.98); TEL/HL 0.14–0.21 (0.18). Distribution. China: Heilongjiang, Jilin, Liaoning. Nanjing, Shanghai, Zhejiang; Taiwan; Vietnam; Laos; Korea; Japan: Tokara Isls. (Nakano-shima), Okinawa-jima, Ishigaki-jima, Iriomote-jima. Remarks. This is the first record of this species from Japan. The body size of the specimens collected from the low-latitude area (e.g. Taiwan, Vietnam, and Japan) is somewhat larger than those from the high-latitude area (e. g. China and Korea).