Tabiboka milleri Henrard, Griswold & Jocqué, 2024, gen. et sp. nov.

Tabiboka milleri gen. et sp. nov. urn:lsid:zoobank.org:act: 33C8C4F1-A241-41E2-9B38-1D31AFBF4BE4 Figs 3–7, 26 Zorocratidae gen. Talatakely sp. Talatakely – Morphbank 2023: images [476092–509952] SpiderAToL, March 3, 2008, https://www.morphbank.net/?id=218853 [accessed 20 Oct. 2023]. Zorodictyna sp. 2 – Polotow et al. 2015. Zorodictyna sp. CG300 – Wheeler et al. 2017. Diagnosis Males of Tabiboka milleri gen. et sp. nov. may be distinguished by the morphology of the palpal tibia with the RTA having an elongate curved spur arising retrobasally and extending distally to the apex of the tibia (Figs 5B–C, 6A–C) (vs T. milloti gen. et sp. nov. and T. polotowae gen. et sp. nov. have a short and stout RTA, Figs 9B–C, 11B–C), a short, broad embolus (E) and a small, slender paraconductor (Pc) largely hidden behind the embolus (Figs 5A–C, 6D–F) (vs in T. milloti and T. polotowae, the E is slender and the Pc is situated between the E and the C, Figs 8F, 9B, 11B, 12E). Females of T. milleri have the epigyne with a broad, swollen, oval median lobe (ML) that extends posteriorly beyond the epigastric furrow, narrow anteriorly (anterior width less than half the total width, Figs 4B, 5D, 7A) (vs in T. milloti, the ML is broad anteriorly, Fig. 9D; and in T. polotowae, the ML is medially narrowed, Figs 11D, 12G). Etymology The species name is a patronym in honor of Jeremy Miller, who collected widely in Madagascar and led a team that carried out a pioneering study of entelegyne spider phylogeny (Miller et al. 2010). Type material Holotype MADAGASCAR – Fianarantsoa Province • ♂; Ranomafana National Park, Talatakely; 21°15′ S, 47°25′ E; 900 m a.s.l.; 5–7 Dec. 1993; C. Griswold, N. Scharff, S. Larcher and R. Andriamasamanana leg.; CASENT_9016094; CAS. Paratypes MADAGASCAR – Fianarantsoa Province • 1 ♀; same collection data as for holotype; [Standard view preparations and images for AToL-Spiders project; SEM preparations Box 79-8, 10, 11, 19, Box 81, 1, 2]; CASENT_9016094; CAS • 1 ♂, 1 ♀; Ranomafana N.P.; 21°12′ S, 47°27′ E; 18 May 1992; V. and B. Roth, S. Kariko leg.; general collecting; [habitus photos, stacked images, H.M. Wood; Apr. 2005]; CASENT_9016098; CAS • 1♂, 1 ♀; same data as for preceding; [♂ palp drawn]; CASENT_9016086; CAS • 1 ♂, 1 ♀; Parc National Ranomafana, Talatekely forest, 42.3 km 58° NE of Fianarantsoa; 21°15′28.0″ S, 47°25′21.8″ E; 1050 m a.s.l.; 24 Dec. 2005 – 14 Jan. 2006; H. Wood, J. Miller, J.J. Rafonomezantsoa, E. Rajeriarison, V. Andriamananony leg.; montane rainforest; general collecting; [AToL DNA Voucher ARACG0300] [HW009/RNAlater: Mad-052]; CASENT_9024004; CAS • 1 ♂; Ranomafana N.P.; 21°12′ S, 47°27′ E; 18 May 1992; V. and B. Roth, S. Kariko leg.; general collecting; [stacked images, HMW, April-2005]; CASENT_9016097; CAS • 1 ♀; Parc National Ranomafana, Vohiparara, 3.6 km W of Ranomafana; 21°14.243′ S, 47°23.842′ E; 1150 m a.s.l.; 13–14 Jan. 2009; D. Andriamalala, C. Griswold, G. Hormiga, A. Saucedo, N. Scharff and H. Wood leg.; primary montane rainforest; general collecting day and night; [NS Photo voucher 893-897; GH photos 5659-5666]; CASENT_9029871; CAS. – Mahajanga Province • 1 ♀; Ampijoroa, National Park Ankarafantsika, near Lake Ravelobe; 16°17′48″ S, 46°48′50″ E; 26–28 Jan. 2009; C. Griswold, A. Saucedo, H. Wood leg.; [SEM female epigyne + used for DNA voucher DP-2014c - “ Zorodictyna sp. 2 ”]; CASENT_9035866; CAS. Other material examined MADAGASCAR – Antananarivo Province • 1 ♂; Tsinjoarivo Forest, Mahatsinjo; 19°40′47.496″ S, 47°46′22.835″ E; 1654 m a.s.l.; 26–29 Aug. 2014; B.L. Fisher leg.; montane rainforest; BLF35410; CASENT_9065708; CAS • 1 ♂; same data as for preceding; CASENT_9065709; CAS • 1 ♂; same data as for preceding; CASENT_9065678; CAS • 1 ♂; same data as for preceding; CASENT_9065579; CAS • 4 ♂♂, 5 ♀♀; same data as for preceding; CASENT_9017817; CAS • 2 ♀♀; Tsinjoarivo Forest, Ankadivory; 19°43′00″ S, 47°49′18″ E; 1369 m a.s.l.; 28 Apr.–1 May 2014; B.L. Fisher et al. leg.; montane rainforest, sifted litter; BLF33370; CASENT_9064661; CAS. – Fianarantsoa Province • 1 ♀; Forêt d’Ankazomivady, 28 km SSW of Ambositra, 4.7 km SW of Ambalamanakana; 20°47′ S, 47°10′ E; 1670 m a.s.l.; 7–14 Jan. 1998; S. Goodman, SG0001 leg.; disturbed upper montane forest; pitfall traps #1–3; CASENT_9016064; FMNH • 9 ♀♀; same data as for preceding; CASENT_9016065; FMNH • 1 ♀; same data as for preceding; CASENT_9016066; FMNH • 9 ♀♀; same data as for preceding; CASENT_9016067; FMNH • 1 ♂; same data as for preceding; CASENT_9016068; FMNH • 1 ♂; Ankazomivady, 29 km SSW of Ambositra; 20°47′ S, 47°10′ E; 1700 m a.s.l.; 7 Jan. 1998; B.L. Fisher leg.; forest; EBD10 pitfall trap; BLF1592b; CASENT_9016087; CAS • 1 ♀; Ambatofitorahana, 33 km S of Ambositra; 20°49′00.00″ S, 47°10′59.99″ E; 13 Mar. 1994; A. Pauly leg.; natural forest; BE_ RMCA_ARA.Ara.201626; RMCA • 24 ♂♂, 5 ♀♀; Ranomafana National Park, Talatakely; 21°12′ S, 47°27′ E (coordinates approximate); 900 m a.s.l.; 4–16 Jan. 2001; D.H. and K.M. Kavanaugh, R.L. Brett, E. Elsom and F. Vargas leg.; mixed tropical forest; pitfall traps #4–16; COL-DHK-2001-001TN; CASENT_9003432; CAS • 1 ♀; same data as for preceding; CASENT_9003534; CAS • 1 ♀; Parc National Ranomafana, Talatakely; 21°15′ S, 47°26′ E; 13–27 Apr. 1998; C.E. Griswold, D.H. Kavanaugh, N.D. Penny, M.J. Raherilalao, J.S. Ranorianarisoa, J. Schweikert and D. Ubick leg.; pitfall traps; CASENT_9006115; CAS • 1 ♂; same data as for preceding; CASENT_9016084; CAS • 1 ♂; Parc National Ranomafana, Talatakely; 21°15′ S, 47°26′ E; 19–30 Apr. 1998, C.E. Griswold, D.H. Kavanaugh, N.D. Penny, M.J. Raherilalao, J.S. Ranorianarisoa, J. Schweikert, D. Ubick leg.; general collecting; CASENT_9016082; CAS • 2 ♀♀; same data as for preceding; CASENT_9016088; CAS • 6 ♂♂; Parc National Ranomafana, Talatakely, trail F; 21°16′ S, 47°25′ E; 940–985 m a.s.l.; 1–21 Nov. 1998; V.F. Lee and K.J. Ribardo leg.; pitfall trap; CASENT_9016089; CAS • 2 ♂♂; same data as for preceding; CASENT_9016090; CAS • 1 ♂; same data as for preceding; CASENT_9016093; CAS • 2 ♀♀; Parc National Ranomafana, Vatoharanana; 21°17′ S, 47°26′ E; 1200 m a.s.l.; 15–29 Apr. 1998; C.E. Griswold, D.H. Kavanaugh, N.D. Penny, M.J. Raherilalao, J.S. Ranorianarisoa, J. Schweikert, D. Ubick leg.; primary forest; CEG-MA-1998-03; CASENT_9016080; CAS • 1 ♀; Parc National Ranomafana, Vatoharanana, trail K; 21°17′ S, 47°26′ E; 990 m a.s.l.; 14–19 Nov. 1998; V.F. Lee and K.J. Ribardo leg.; pitfall trap; VFL-KJR-02; CASENT_9016091; CAS • 1 ♀; same data as for preceding; CASENT_9016092; CAS • 1 ♀; Parc National Ranomafana, Vatoharanana River, 4.1 km 231° SW of Ranomafana; 21°17′24″ S, 47°26′00″ E; 1100 m a.s.l.; 27–31 Mar. 2003; Fisher-Griswold Arthropod Team leg.; montane rainforest; general day collecting; BLF8394; CASENT_9017837; CAS • 1 ♀; Parc National Ranomafana, Vohiparara Piste Touristique; 21°14′ S, 47°24′ E; 1000 m a.s.l.; 23 Apr. 1998; C.E. Griswold, D.H. Kavanaugh, N.D. Penny, M.J. Raherilalao, E. Rajeriarison, J.S. Ranorianarisoa, J. Schweikert, D. Ubick leg.; running in litter; general collecting; CEG-MA-1998; CASENT_9016081; CAS • 6 ♂♂; Ranomafana National Park, Vohiparara; 21°12′ S, 47°27′ E (coordinates approximate); 1150 m a.s.l.; 2–22 Jan. 2001; D.H. and K.M. Kavanaugh, R.L. Brett, E. Elsom, F. Vargas leg.; mixed tropical forest; general collecting; COL- DHK-2001-004; CASENT_9003479; CAS • 6 ♀♀; Parc National Ranomafana, 2.3 km N of Vohiparara Village; 21°13′ S, 47°23′ E; 1100 m a.s.l.; 18 Apr. 1998; C.E. Griswold, D.H. Kavanaugh, N.D. Penny, M.J. Raherilalao, E. Rajeriarison, J.S. Ranorianarisoa, J. Schweikert, D. Ubick leg.; CASENT_9016083; CAS • 6♀♀; same data as for preceding; CASENT_9016085; CAS • 2 ♀♀; Ranomafana N.P., Trail F; 21°12′ S, 47°27′ E (coordinates approximate); 18 May 1992; Albert for Kariko, Roth leg.; sifting #1; CASENT_9030889; MCZ • 1 ♀; Ranomafana N.P., Maharira, summit; 21°12′ S, 47°27′ E; 9 Apr. 1992; Albert for Kariko, Roth leg.; cryptic #2; CASENT_9030890; MCZ • 1 ♂; Ranomafana N.P., Trail M; 21°12′ S, 47°27′ E (coordinates approximate); 15 May 1992; Albert for Kariko, Roth leg.; sifting 1hr. samples #2; CASENT_9030891; MCZ • 1 ♀; Ranomafana N.P., Trail M; 21°12′ S, 47°27′ E (coordinates approximate); 15 May 1992; Albert for Kariko, Roth leg.; sifting; CASENT_9030892; MCZ • 3 ♀♀; Ranomafana N.P., Trail B; 21°12′ S, 47°27′ E (coordinates approximate); 825 m a.s.l.; 19 May 1992; Albert for Kariko, Roth leg.; sifting, #1; CASENT_9030893; MCZ • 1 ♀; Ranomafana N.P., Maharira, summit; 21°12′ S, 47°27′ E (coordinates approximate); 11 Apr 1992; Emile for Kariko, Roth leg.; sifting #3; CASENT_9030894; MCZ • 2 ♀♀; Ranomafana N.P.; 21°12′ S, 47°27′ E; Apr. 1992; V. and B. Roth, S. Kariko leg.; forest; general collecting from foliage; Ranomafana-1000-92; CASENT_9016099; CAS • 1 ♂, 1 ♀; Ranomafana N.P., trail F, secondary forest; 21°12′ S, 47°27′ E (coordinates approximate); Apr. 1992; Albert for Kariko, Roth leg.; sifting #3; MCZ-VBR-0116; CASENT_9016100; MCZ • 4 ♂♂, 7 ♀♀; Ranomafana N.P.; 21°12′ S, 47°27′ E (coordinates approximate); 20–31 May 1992; V. and B. Roth leg.; can traps; MCZ-VBR-02; CASENT_9016101; MCZ • 1 ♂; same data as for preceding; CASENT_9016102; CAS. Description Male holotype (CASENT_9016094; CAS) MEASUREMENTS. Total length: 10.50. Carapace: length 6.10, width 4.40, height 2.20. COLORATION (Fig. 3A–C, G). Carapace brownish yellow, darker along margin, with two faint, longitudinal dark bands along margin of pars cephalica; black on ocular area and surrounding all eyes; chelicerae brownish yellow; sternum yellow, unmarked; legs brownish yellow, unmarked; abdomen brownish yellow with darker cardiac mark and dorsolateral dark bands that break up into chevrons posteriorly, sides brownish yellow mottled with dark marks, venter yellow except darker around spinneret base. STERNUM (Fig. 3B). Oval with slightly sinuous margins; 2.30 wide, 2.60 long. CHILUM. 0.13 high, 0.35 wide. EYES. ALE 0.18; AME 0.13; PLE 0.18; PME 0.15; ALE–AME 0