Euura hebes ZMUO.

203 Euura hebes (Konow, 1907) Figs 230, 454 Amauronematus hebes Konow, 1907: 17–18. Lectotype designated by Muche (1975). Amauronematus pristiphorinus Malaise, 1921a: 2. Synonymy by Lindqvist (1941b). Amauronematus pribaicalicus Lindqvist, 1970: 100–102. Syn. nov. Amauronematus pristiphoroides Lindqvist, 1972: 72–73. Syn. nov. Pristiphora albomarginata Lindqvist, 1974a: 117–118. Syn. nov. Amauronematus harpicola Bogacheva, 1977: 87. Lectotype ♀ (ZMMU), designated by Zinovjev & Schmidt (1994), not examined. Type locality: Russia, Polar Urals, Kharp Biological Station. Syn. nov. Diagnosis Mostly black, brown or pale pterostigma, “ Amauronematus - type ” head, bifid claws, square-shaped valvula 3 with distinct invagination. Most similar species is E. aegra (character states in parentheses), from which it differs by having more matt mesepisternum (usually weakly matt) and darker pronotum (usually extensively pale). Males possibly not clearly distinguishable from other aegra group species. Although pribaicalicus, pristiphoroides, and albomarginata have colouration (pronotum and clypeus usually at least partly pale, valvula 3 completely or mostly black) and mesepisternum sculpture (slightly matt) more similar to aegra than hebes (pronotum and clypeus completely or nearly completely black, valvula 3 extensively pale; mesepisternum strongly matt) we consider these nominal taxa as synonyms of hebes based on the saw, host plant, and genetics. Based on the description provided by Zinovjev & Schmidt (1994), colouration of harpicola (reared from Salix) seems overall more similar to pribaicalicus than hebes, but colouration of the sawsheath (valvula 3) fits with typical hebes. Saws of the above-mentioned nominal taxa do not seem to be different. The species is not monophyletic based on current sampling of nuclear genes, but this is the case even for individual females when haplotypes of nuclear genes are separated, indicating incomplete lineage sorting and not necessarily the presence of additional species. Type material examined Lectotype Amauronematus hebes SWEDEN • ♀; Northern Lapland; SDEI, GBIF-GISHym4093. Holotype Amauronematus pristiphorinus SWEDEN – Torne Lappmark • ♀; Torneträsk; 68.22° N, 19.74° E; 400 m a.s.l.; R. Malaise leg.; NHRS, NHRS-HEVA000003841. Holotype Amauronematus pribaicalicus RUSSIA – Irkutskaja oblast • ♀; Irkutsk, Oljhonsk, Popovo; 52.741° N, 106.309° E; 10 Apr. 1968; B.N. Verzhutskij leg.; MZH, http://id.luomus.fi/GL.5128. Holotype Amauronematus pristiphoroides RUSSIA – Kamchatka oblast • ♀; Nischne-Kamtchatsk, Lake Azhabachye; 56.166° N, 161.893° E; R. Malaise leg.; NHRS, NHRS-HEVA000006425. Holotype Pristiphora albomarginata FINLAND – Kittilä Lapland • 1 ♀; Muonio, Pallastunturi; 68.100° N, 23.999° E; 13 Jun. 1967; O. Ranin leg.; MZH, http://id.luomus.fi/GL.3403. Notes A COI sequence (658 bp) of the holotype is most similar to DEI-GISHym19918 (E. hebes). Host plants Salix sp. or spp. (Kontuniemi 1960): at least grey-leaved species are hosts. Genetics COI Based on 19 specimens, maximum within-species distance is 4.1% and the nearest neighbour, diverging by a minimum of 0.15%, is Euura lindqvisti. Nuclear Based on 13 specimens, maximum within-species distance is 0.73% (0.65% based on haplotypes of individual females). The nearest neighbour, diverging by a minimum of 0.18%, is Euura lindqvisti. Distribution and material examined Palaearctic. Specimens studied are from Finland, Russia, and Sweden.