Anthobium atrocephalum

Anthobium atrocephalum (Gyllenhal, 1827) (Figs 1–3, 7–9) Omalium atrocephalum Gyllenhal, 1827: 463. Anthobium ruficolle Stephens, 1834: 340. Anthobium longipenne Stephens, 1834: 342. Lathrimaem cicatrix Mulsant and Rey, 1880: 130. Lathrimaeum concolor Delahon, 1914: 621. Lathrimaem (Lathrimaeum) sardoum Scheerpeltz 1961: 84, 92. Lathrimaeum atrocephalum sardoum: Zanetti 1986: 96. Anthobium atrocephalum: Khachikov 1998: 6, Assing 2004: 676, 2011: 183, Bordoni 2010: 52, Assing & Schülke 2019: 98, Khachikov et al. 2021: 249, Salnitska et al. 2022: 57. Lathrimaeum gracilicorne Luze, 1905: 66 syn. n. Lathrimaeum (Lathrimaeum) gracilicorne: Scheerpeltz, 1961: 83. Lathrimaeum abantense Fagel, 1968: 121 syn. n. Anthobium abantense: Assing 2013: 106, 2014: 1135; Bordoni 2010: 52. Lathrimaeum urgelense Fagel, 1968: 122 syn. n. Other references see in Shavrin (2020). Type material examined. Holotype of Lathrimaeum gracilicorne Luze, 1905 (Fig. 1; dissected; left side of the pronotum is slightly damaged): ‘longicorne Fv. | Hammam R`iza | Africa bor. | leg. Hauser’ <handwritten>, ‘c. Eppelsh. | Steind. d.’ <printed>, ‘Type. gracili - | corne Luze’ <handwritten>, ‘atrocephalum | Gyll. var.’ <handwritten>, ‘TYPUS’ <red, printed>, ‘ Anthobium | atrocephalum (Gyllenhal, 1827) | Shavrin A.V. det. 2024’ <printed> (NMW). Holotype of Lathrimaeum abantense Fagel, 1968 ♂ (Fig. 2; dissected prior to the present study): [red round label], ‘Anatolie occid. | Abant Gölü | 1450 m – V.1967 | G. Fagel’ <printed>, ‘TYPE’ <red printed label with black margin>, [blue round label], ‘G. Fagel det., [printed] | abantense | n. sp.’ <handwritten>, ‘ Anthobium | atrocephalum (Gyllenhal, 1827) | Shavrin A.V. det. 2024’ <printed> (IRSNB). Paratype ♂ (dissected prior to the present study): [red round label], ‘Anatolie occid. | Abant Dagh. | 1400-1550 m | V.1967 G.Fagel’ <printed>, ‘G. Fagel det. [printed] | sp. apud | atrocephalum | Gyll.’ <handwritten>, ‘G. Fagel det., [printed] | abantense | n. sp.’ <handwritten>, ‘ Para- | type’ <red printed label with black margin>, ‘ Anthobium | atrocephalum (Gyllenhal, 1827) | Shavrin A.V. det. 2024’ <printed> (IRSNB). Holotype of Anthobium urgelense Fagel, 1968 ♂ (Fig. 3; dissected prior to the present study): ‘ Catalogne: Seo de | Urgel: Sierra del Cadi | 1000 m, V-VI-1962 | G.Fagel 104 [handwritten]’ <printed>, [red small round label], ‘G.Fagel det., 1969 [69 - handwritten] | L.atrocephalum | ?? [handwritten] Gyll.’ <printed>, ‘G. Fagel det., | urgelense [handwritten] | n.sp [handwritten]’ <printed>, ‘TYPE’ <red printed label with black margin>, [blue small round label], ‘ Anthobium | atrocephalum (Gyllenhal, 1827) | Shavrin A.V. det. 2024’ <printed> (IRSNB). Paratypes (all specimens dissected prior to the present study): 1 ♂: [red small round label], ‘ Catalogne: Seo de | Urgel: Sierra del Cadi | 1000 m, V-VI-1962 | G.Fagel 109’ [handwritten] <printed>, ‘G. Fagel det., | urgelense [handwritten] | n.sp [handwritten]’ <printed>, ‘ Para- | type’ <red printed label with black margin>, [blue small round label] (IRSNB); 1 ♂: [red small round label], ‘ Espagne | Castille | Puerto de Bejar | 800-1400 m | V – VI 1957 G.Fagel’ <printed>, ‘G. Fagel det., | urgelense [handwritten] | n.sp [handwritten]’ <printed>, ‘ Para- | type’ <red printed label with black margin>, [blue small round label] (IRSNB); 1 ♀: labels as previous except first printed label: ‘ ♀ ’ (IRSNB). All paratypes with additional printed label: ‘ Anthobium | atrocephalum (Gyllenhal, 1827) | Shavrin A.V. det. 2024’. Additional material examined. MOROCCO: 2 ♀♀: Azrou. 19- 20.05.1999. R. Milan leg. (NMPC); ALGERIA: 21 ♂♂, 15 ♀♀: Gde Kabylie, Forêt d`Akfadou, 8 km E Adekar. 1300 m a.s.l. 15.05.1988. C. Besuchet, I. Löbl & D. Burckhardt leg. (MHNG, cSh); 2 ♂♂, 1 ♀: same data, 1200 m, 16.05.1988 (MHNG); 1 ♂, 2 ♀♀: same data, 1050 m a.s.l. (MHNG); 1 ♀: same data, 9 km W Adekar. 1300 m a.s.l. 17.05.1988 (MHNG); 1 ♀: ‘Massif des Mouzala’ (NMPC); PORTUGAL: 1 ♂: Arrabães, Villa Real. 600 m a.s.l. 27.07.1981. I. Löbl leg. (MHNG); SPAIN: HUESCA: 1 ♂: Valle de Ordesa, 12.06.1966. C. Besuchet leg. (MHNG); TERUEL: 2 ♂♂, 3 ♀♀: Griegas. 1500 m a.s.l. 21.05.1960. C. Besuchet leg. (MHNG); 1 ♀: Entre Alcala de la Selva et Gudar. 1400 m a.s.l. 22.05.1960. C. Besuchet leg. (MHNG); 1 ♀: Noguera. 1550 m a.s.l. 20.05.1960. C. Besuchet leg. (MHNG); FRANCE: ROUSSILON: 1 ♂, 3 ♀♀: Montferrer-Corsavy. 14.04.1977. I. Löbl leg. (MHNG); 1 ♂: Parcigoule. 15.4.1977. I. Löbl leg. (MNHG); 2 ♂♂, 2 ♀♀: same data, Le Tech. (MHNG); FOIX: 1 ♀: Crouzette. 08.1967 (MHNG); DAUPHINÉ: 5 ♀♀: Vercors, Col de Limonches. 1000 m a.s.l. 09.04.1980. I. Löbl leg. (MHNG); SAVOIE: 1 ♂: Col du Chat. 24.04.1975. I. Löbl leg. (MHNG); 1 ♀: Le Bourget du Lac. 24.04.1975. I. Löbl leg. (MHNG); PROVENCE: 4 ♀♀: Uvernet. 1250 m a.s.l. 05.06.1974. I. Löbl leg. (MHNG); 3 ♀♀: Ubaye le Martinet. 03.06.1974. I. Löbl leg. (MHNG); 1 ♂, 1 ♀: Cousson. 05.06.1970. Raffaldi leg. (MHNG); 1 ♂: Gard, Mt. Aigonal. 1450 m a.s.l. 06.05.1980. I. Löbl leg. (MHNG); 2 ♀♀: Seranon. 22.07.1968 (MHNG); 1 ♀: Col des Champs. 07.1965. (MHNG); NICE: 1 ♀: Sospel. 10.1962 (MHNG); CORSICA: 1 ♂: Vizzavona. 1000 m a.s.l. 18.07.1978. I. Löbl leg. (MHNG); ITALY: PIEMONTE: 1 ♀: Crissolo. 11.07.1965 (MHNG); VENETO: 1 ♂: Mt. Baldo Ferrara. 880 m a.s.l. 31.05.1977. I. Löbl leg. (MHNG); 1 ♂: Monte Grappa. 1800 m a.s.l. 09.1969. Curti leg. (MHNG); 2 ♂♂, 3 ♀♀: Fosse. 20.06.1968 (MHNG); EMILIA-ROMAGNA: 2 ♂♂, 6 ♀♀: Grotte Santa Maria Madalena, Valestra. 12.06.1976. I. Löbl leg. (MHNG); SARDEGNA: 1 ♀: Mt. Spada. 1100 m a.s.l. 14.4.1977. S. Vit leg. (MHNG); SERBIA: 10 ♂♂, 7 ♀♀: Krupanj. 450 m a.s.l. 24.05.1984. C.Besuchet leg. (MHNG); 6 ♂♂, 6 ♀♀: Rtanj Planina. 600 m a.s.l. 27.05.1984. C.Besuchet leg. (MHNG); 1 ♂, 2 ♀♀: Tara Planina, Mitrovac. 900 m a.s.l. 23.05.1984. C.Besuchet leg. (MHNG); 1 ♂: same data, 950 m a.s.l. (MHNG); 2 ♀♀: Stol Planina. 900 m a.s.l. 26.05.1984. C.Besuchet leg. (MHNG); UKRAINE: CRIMEA: 2 ex.: Ay Petri (ZIN); 1 ex.: ‘Tauria, Пеньдикюль [Pendikyul`]. 11.4.14 V. Pliginski’ (ZIN); 3 ex: Roman-Kosh. 22.05.1948. K. Arnol`di leg. (ZIN); RUSSIA: STAVROPOL TERRITORY: 2 ♀♀: Zheleznovodsk, Razvalka Mt. 8.06.1985. A.B. Ryvkin leg. (cR); KRASNODAR TERRITORY: 1 ♂, 3 ♀♀: environs of Krasnodar. Floodplain forest, litter. 14.04.1995. A. Solodovnikov leg. (ZMUC); 2 ♂♂, 1 ♀: Goryachiy Klyuch, gorge, north slope. 08.04.1952. K. Arnol`di leg. (cR., ZMM); ADYGEA: 1 ♂: Lagonaki. 1200 m a.s.l. 14.06.1990. Ochrimenko leg. (SMNS); 1 ♀: Abadzekhskaya. 25.09.1986. N.D. Silichenkov leg. (cSh); KARACHAY-CHERKESSIA: 1 ♂: Dombay. 2500 m a.s.l. 10.10.1983. V. Biža leg. (cH); 2 ♀♀: Teberda, Dombay. 07.2010. E.A. Khachikov leg. (cKh); 2 ♂: Teberda State Nature Biosphere Reserve, Malaya Khatipara Mt. 1700 m a.s.l. 30.05.1985. A.B. Ryvkin leg. (cR); 2 ♂♂, 4 ♀♀: same Reserve. 1.06.1985. A.B. Ryvkin leg. (cR, cSh); 1 ♂: same Reserve, Kizgich Canyon near Arkhyz. 1400 m a.s.l., swamp with Carex spp., Juncus sp. 05.06.1985. A.B. Ryvkin leg. (cR); 5 ♂♂, 9 ♀♀: same Reserve, gorge of Gonachkhir River. 1700-1900 m a.s.l., forest with Picea, Betula and Fagus, in litter. 01.06.1985. S.I. Golovatch & A.B. Ryvkin leg. (cR, cSh); NORTH OSSETIA-ALANIA: 2 ♀♀: Sunzhenskiy Mts., between Kardzhin and El`khotovo villages. 570 m a.s.l., NW slope, Fagus forest with Rubus and Festuca. 03.10.1985. S.K. Alekseev leg. (cR); 2 ♂♂: Ardon River, Zintsar. 13.04.1984. Forest swamp. S.K. Alekseev leg. (cR); 1 ♂: valley of Ardok River. 26.08.1985. S.K. Alekseev leg. (cSh); DAGESTAN: 3 ♂♂, 1 ♀: Samur Valley, Garakh ca. 35 km SW of Magaramkent. 700-800 m a.s.l., Crataegus, Quercus, Acer, Rosa etc., scrub litter. 23.10.1987. S.I. Golovatch leg. (cSh); 3 ♂♂, 3 ♀♀: Dagestan. 1987. S.I. Golovatch leg. (cSh); ARMENIA: LORI: 11 ♂♂, 25 ♀♀: Stepanavan. 1500-1650 m a.s.l., forest with Quercus, Fagus, Carpinus etc., litter and under bark. 21- 22.05.1987. S.I. Golovatch & K.Yu. Es`kov leg. (cSh, ZMM); 1 ♂, 1 ♀: Odzun W of Alaverdi. 1500-1550 m a.s.l., forest with Quercus, Fagus, Carpinus, litter and under stones, 23- 24.07.1987. S.I. Golovatch & K.Yu. Es`kov leg. (ZMM); 2 ♂: Bazum Mt. Ridge, N slope Puskiri Pass. 1700 m a.s.l., forest with Fagus and Pinus, litter. 22.05.1987. S.Golovatch & K.Yu. Es`kov leg. (cSh); TAVUSH: 5 ♂♂, 7 ♀♀: Dilijan National Park, Aknabat Valley near Salakh. 1400-1500 m a.s.l., forest with Fagus, Taxus, Carpinus etc., in litter, under bark and stones. 17.04.1983. S.I. Golovatch leg. (cSh, ZMM); 3 ♀♀: same Park, Agartsyn. 1350-1400 m a.s.l., forest with Fagus and Acer, litter. 28- 29.05.1987. S.I. Golovatch & K. Eskov leg. (cSh); 1 ♂, 3 ♀♀: W of Shamshadyn halfway between Idjevan and Berd. 1500-1600 m a.s.l. 26- 27.05.1987. S.I. Golovatch & K.Yu. Es`kov leg. (cSh); 5 ♂♂, 3 ♀: Idjevan District, Tsakhkavan. 850-900 m, forest with Quercus, Acer, Carpinus etc., litter and tree hole. 25.07.1987. S.I. Golovatch & K.Yu. Es`kov leg. (cSh); ARARAT: 19 ♂♂, 21 ♀♀: N Vartashen. 900 m a.s.l., forest with Fagus and Quercus, in litter. 02.05.1987. S.I. Golovatch & K.Yu. Es`kov leg. (cSh, ZMM); SYUNIK: 2 ♂♂, 1 ♀: ca. 4 km NNW Megri, village Lehvaz. 1000 m a.s.l., Juglans and oak shrub with Polyurus and Rosa, litter and under stones. 24- 25.04.1983. S.I. Golovatch leg. (cSh, ZMM); 4 ♂♂, 1 ♀: Kapan District, Shikahogh State Reserve, vill. Shikahogh. 900-950 m a.s.l., forest with Quercus, Fagus and Carpinus, litter and under stones. 28.04.1983. S.I. Golovatch leg. (cSh); 1 ♂: Nerkin Hand. Plane-tree grove. 30.04.1983. A.B. Ryvkin leg. (cR); GEORGIA: ABKHAZIA: 1 ♂: W Gudauta, Myussera State Reserve. 100 m a.s.l., in leaf litter on slope. 10.04.1983. S.I. Golovatch leg. (ZMM); 1 ♂: Sukhumi, mouth of Kelasuri River. 16.09.1976. A.B. Ryvkin leg. (cR); KAKHETI: 1 ♀: Lagodekh Nature Reserve. 500 m a.s.l., wet forest litter near water. 06.05.1983. S.I. Golovatch leg. (ZMM); 2 ♀♀: Batsara Strict State Reserve, ca 20 km N of Akhmeta. 800-850 m a.s.l., forest with Fagus and Castanea, litter. 05- 06.05.1987. S.I. Golovatch &