Aname L. Koch 1873

Genus Aname L. Koch, 1873 Aname L. Koch, 1873: 465. Dekana Hogg, 1902: 138 (synonymised by Raven 1981: 328). Dolichosternum Rainbow & Pulleine, 1918: 168 (synonymised by Raven 1981: 328). Sungenia Rainbow & Pulleine, 1918: 162 (synonymised by Raven 1981: 328). Type species Aname: Aname pallida L. Koch, 1873, by monotypy. Dekana: Dekana diversicolor Hogg, 1902, by original designation. Dolichosternum: Dolichosternum attenuatum Rainbow & Pulleine, 1918, by monotypy. Sungenia: Chenistonia atra Strand, 1913, by monotypy. Diagnosis Modified from Harvey et al. (2018): species of Aname can be diagnosed from all other Australian anamid genera by the presence of a ventral asetose depression on the palp tibia of males (see Fig. 2I and Figs 3–4 for further examples). They can be further distinguished from Chenistonia Hogg, 1901, Proshermacha Simon, 1908, Teyloides Main, 1985, Teyl Main, 1975, and Namea Raven, 1984 (all members of the subfamily Teylinae) by the presence of a short cymbium without a medial constriction, and further from Namea and Teyl by the presence of a tibial megaspur. Both adult males and females can further be distinguished from Chenistonia, Proshermacha, Teyloides, Teyl, and Namea by the presence of a relatively broad (extending laterally and anteriorly) patch of cuspules on the maxillae. Description See Harvey et al. (2018). Key to the species complexes of Aname from tropical and subtropical eastern Australia, including miscellaneous species Note that the keys in this paper use Boolean operators, with character states separated by a comma and linked by an “or” operator denoting states that may or may not apply (and only one must apply for the requirements of the statement to be met). Semi-colons separate implicit relevant states and should be treated as “and” operators in a Boolean context (with any exceptions noted in brackets). 1. Males.................................................................................................................................................. 2 – Females ............................................................................................................................................ 27 Male morphology 2. Embolus length>1.5 × bulb length (e.g., Figs 3A–D, E [x], F; 4D, F–H, L–M, Q–R)....................... 3 – Embolus shorter (e.g., Figs 3E [ix], G; 4A–C, E, I–K, N–P).............................................................. 16 3. Carapace length <4.0 mm; embolus long, straight, and reflexed relative to the bulb (e.g., Fig. 4D). .......................................................................................................................... flexicaudula -complex – Carapace longer ................................................................................................................................. 4 4. Copulatory organ with a highly reflexed and broad corkscrew-shaped embolus (Fig. 4Q) ................ .......................................................................................................................... A. serpentina sp. nov. – Copulatory organ otherwise............................................................................................................... 5 5. Palp tibia with a small, tight patch of thorn-like setae on retrolateral side of asetose depression; metatarsus I proximal excavation length ~0.6 × metatarsus I length, with a sharp heel (Fig. 4J)...... ....................................................................................................................... A. camara Raven, 1985 – Palp tibia without tight patch of thorn-like setae; metatarsus I with a shorter proximal excavation, or a more rounded heel........................................................................................................................... 6 6. Metatarsus I proximal excavation length <0.5 × metatarsus I length, with a prominent and sharp heel, and a relatively straight distal pad (e.g., Fig. 3A)....................................................................... .................................................................................... pallida -complex (except A. platensis sp. nov.) – Metatarsus I otherwise....................................................................................................................... 7 7. Embolus with a relatively wide basal section that tapers to a curvy and sinuous distal section after about 0.4–0.5 of length (e.g., Fig. 3B)................................................................. eddieorum -complex – Embolus otherwise............................................................................................................................. 8 8. Embolus thin, with a sharp bend at about 0.2–0.4 of length, before a relatively straight distal section (e.g., Fig. 3D).................................................................................................... robertsorum -complex – Embolus otherwise............................................................................................................................. 9 9. Tibial megaspine length <0.2 × tibia I length, usually angled almost parallel with tibia; prolateral patellae of pedipalp and leg I with three or more spines (or spine ‘sockets’ if spines have been lost); sternum length <1.2 × width, covered in short setae (e.g., Fig. 3F)....................... warialda -complex – Tibial megaspine longer, or prolateral patellae of pedipalp and leg I and pedipalp with two spines or less, or sternum narrower................................................................................................................. 10 10. Palp tibia asetose depression length ~ 0.8 × palp tibia length; embolus length>2.5 × bulb length (Fig. 4G, M)......................................................................................................................................11 – Asetose depression or embolus shorter............................................................................................ 12 11. Embolus length ~4.6 × bulb length (Fig. 4G) ....................................................... A. distorta sp. nov. – Embolus shorter (~2.8 × bulb length) (Fig. 4K) ................................................. A. consuelo sp. nov. 12. Metatarsus I proximal excavation>0.5 × metatarsus I length; copulatory organ with bulb tapering into the embolus, a strong curve at about 0.6 of length, and a small hook at the tip (Fig. 4R)............................................................................................................. A. viridiensis sp. nov. – Proximal excavation shorter; copulatory organ otherwise .............................................................. 13 13. Copulatory organ with the bulb tapering into the embolus, the embolus thicker at the base before tapering and curving sharply at about 0.6 of its length (Fig. 3E)............... barrema -complex, in part – Copulatory organ otherwise............................................................................................................. 14 14. Metatarsus I length ~ 3.5 × width (Fig. 4F) ........................... A. platensis sp. nov. (pallida -complex) – Metatarsus I thinner (length> 3.9 × width) ..................................................................................... 15 15. Sternum length>1.25 × width; tibial spur triangular (e.g., Fig. 3C) .................... barakula -complex – Sternum broader; tibial spur more digitiform (Fig. 4H) .................................. A. lawrenceae sp. nov. 16. Carapace length <4.0 mm ............................................................................................................... 17 – Carapace longer ............................................................................................................................... 19 17. Copulatory organ with bulb tapering into embolus, embolus with thick base; palp tibia with thorn-like setae along retrolateral edge of asetose depression, getting denser proximally (Fig. 4M)................................................................................................................ A. litoralis sp. nov. – Copulatory organ with more demarcated bulb and embolus; palp tibia without thorn-like setae (e.g., Fig. 4C, E)........................................................................................................................................ 18 18. Embolus with sharp tip; metatarsus I length> 4 × width (e.g., Fig. 4C)................. mariala -complex – Embolus with wider tip; metatarsus I thicker (e.g., Fig. 4E)............................... savannella -complex 19. Embolus thick and relatively straight, with a slight angle change at the tip; tibial spur triangular (e.g., Fig. 3G)..................................................... rubrochelicera -complex (except A. savannensis sp. nov.) – Copulatory organ otherwise, or tibia I with a more digitiform spur................................................ 20 20. Copulatory organ with an angular bulb with a slight ridge adjacent to the embolus (e.g., Fig. 4A)............................................................................................................. callitra -complex – Copulatory organ otherwise............................................................................................................. 21 21. Tibial spur digitiform (e.g., Fig. 4I, N–O)....................................................................................... 22 – Tibial spur more triangular (e.g., Fig. 4B)....................................................................................... 24 22. Tibial megaspine> 0.3 × tibia I length; embolus straight, with slight bend near tip (Fig. 4O) ............................................................................................................. A. ethabuka sp. nov. – Tibial megaspine shorter; embolus more strongly curved (Fig. 4I, N)...........................................