Anillini Jeannel 1937

Tribe Anillini Jeannel, 1937 Adult diagnosis. In the United States, adult specimens of Anillini are the only carabids that are both eyeless and possess subulate palpomeres. Larval diagnosis. Late-instar larvae of Anillini in South Carolina share the following characters: body pale and soft, largely unsclerotized (Fig. 13 A, B); legs with single tarsal claw; stemmata absent; coronal suture absent; retinaculum large; penicillus present and consisting of several setae; urogomphus with seven large setae (Fig. 14 A, E). Other known carabid larvae in SC of similar size with a single tarsal claw and potentially lacking stemmata are the genera Trechus, Semiardistomis, Clivina, and the tribe Tachyini. Larvae of Trechus are easily recognized by the possession of 4 - segmented labial palps and 5 - segmented maxillary palps (versus 2 - and 4 - segmented, respectively, in anillines). Semiardistomis and Clivina can be readily separated by the small size of the retinaculum and a penicillus consisting of a single, large seta. South Carolina anilline larvae are similar to larvae of the tribe Tachyini, but differ by having seven large setae on the urogomphus (Fig. 14 E), versus six in tachyines.