Cephennodes dilatatus Jałoszyński, 2025, sp. nov.

Cephennodes (s. str.) dilatatus sp. nov. (Figs 165–168) Type material. Holotype: CHINA (YUNNAN Prov.): ♂, two labels: “CHINA mer.: Yunnan prov. / (pass SW from Baoshan / Gaoligong Shan) / 4. - 8.6.2005 / sifted from base of old tree / Oto Nakladal lgt.” [white, printed]; “ CEPHENNODES (s. str.) / dilatatus m. / P. Jałoszyński, 2025 / HOLOTYPUS ” [red, printed] (MNHW). Diagnosis. Body small, BL around 1.2 mm; antennae gradually thickened; punctures on clypeus and frontovertexal region very small but sharply marked and dense, separated by spaces subequal to their diameters, pronotum with slightly larger, more distinct and similarly dense punctures, elytra with punctures similar to those on head; subhumeral lines as long as about 0.2 × EL. Male: abdomen (Fig. 166) with sternite V modified, bearing conspicuously broad median subtriangular transverse carina; metatibia (Fig. 165) strongly thickened in subapical region; aedeagus (Figs 167–168) in ventral view broadest in subapical region, apical projection oblique in relation to long axis of median lobe, its apex not hidden behind apex of capsular region of median lobe but adpressed against it. Description. Body of male (Fig. 165) strongly convex, oval, with indistinct constriction between pronotum and elytra, moderately dark brown with lighter tarsi, covered with light brown setae. BL 1.20 mm. Head broadest across moderately large and moderately strongly convex, coarsely faceted eyes, HL 0.18 mm, HW 0.28 mm; frontovertexal region weakly convex at middle; supraantennal tubercles weakly elevated. Punctures on clypeus and frontovertexal region very small but sharply marked and dense, separated by spaces subequal to their diameters; setae sparse, short, suberect. Antennae moderately long and slender, gradually thickened, AnL 0.60 mm, antennomere 1 about as long as broad, 2 strongly elongate, 3 distinctly transverse, 4–7 each about as long as broad, 8–10 each weakly transverse, 11 as long as 9 and 10 combined, about 1.8 × as long as broad. Pronotum semi-oval, strongly convex at middle and flattened near hind corners, broadest near middle and distinctly narrowing posterad; PL 0.35 mm, PW 0.50 mm. Anterior margin in strictly dorsal view nearly straight; lateral margins strongly rounded in anterior half and less strongly posteriorly, finely microserrate, posterior corners nearly right-angled; posterior margin shallowly bisinuate; lateral marginal carinae simple (i.e., not doubled); lateral antebasal pits shallow but distinct, each slightly closer to lateral than posterior pronotal margin. Punctures on pronotal disc slightly larger and deeper than those on head, distinct, separated by spaces subequal to their diameters; setae short and moderately dense, suberect. Elytra slightly wider than pronotum, together oval, broadest near anterior third; EL 0.68 mm, EW 0.55 mm, EI 1.23; subhumeral lines as long as about 0.2 × EL, developed as sharp borders between elevated humerus and lower mesal region, strongly divergent posterad; basal elytral fovea on each elytron situated slightly closer to lateral margin of mesoscutellum than to subhumeral line; elytral apices separately rounded. Punctures on elytra similar to those on head, shallower and slightly sparser than those on pronotum; setae similar to those on pronotum. Hind wings not studied. Metaventrite with lateral impressions as long as about half of its length. Legs long and slender, metatibiae with wide subtriangular outer lateral subapical expansion (Fig. 165). Abdomen (Fig. 166) with sternite V bearing long transverse subtriangular median carina. Aedeagus (Figs 167–168) of simonis form, AeL 0.14 mm; median lobe in ventral view broadest in subapical region, apical projection oblique in relation to long axis of median lobe, its apex not hidden behind apex of capsular region of median lobe but adpressed against it; parameres long and slender, one nearly reaching apex of aedeagus, each with one short apical seta. Female. Unknown. Distribution. S China: Yunnan. Etymology. The adjective dilatatus refers to the dilated hind tibiae. Remarks. Among Chinese species, C. dilatatus can be identified by a combination of the following characters: aedeagus in ventral view broadest in subapical region, apical projection oblique in relation to long axis of median lobe, its apex not hidden behind apex of capsular region of median lobe but adpressed against it. Among the Himalayan species, C. dolakhanus, C. manangensis, C. monolaminatus, and C. thakanus have a modified sternite V and unmodified IV, as in C. dilatatus. Cephennodes dolakhanus and C. thakanus have clearly different shapes of the apical plates of the median lobe. Cephennodes manangensis and C. monolaminatus have the aedeagi most similar to that of C. dilatatus. Cephennodes manangensis has a different pattern of punctures on body parts: on the pronotal disc punctures are much smaller and shallower than those on the head, but denser, and on elytra slightly more distinct than those on pronotum, whereas in C. dilatatus the pronotum is covered with punctures slightly larger, more distinct and similarly dense as those on the frontovertexal region, and elytra with punctures similar to those on the head. Cephennodes monolaminatus is distinctly larger than C. dilatatus, body length respectively 1.45–1.58 mm and 1.20 mm, and has the pronotal disc covered with fine and inconspicuous punctures.

Publication Details

Journal
Zenodo (CERN European Organization for Nuclear Research)
Published
2025-07-18
DOI
https://doi.org/10.5281/zenodo.16606590
Primary Topic
Subterranean biodiversity and taxonomy
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article

Cephennodes dilatatus Jałoszyński, 2025, sp. nov.

Zenodo (CERN European Organization for Nuclear Research)
Subterranean biodiversity and taxonomy
article

Cephennodes dilatatus Jałoszyński, 2025, sp. nov.

article en

Abstract

Cephennodes (s. str.) dilatatus sp. nov. (Figs 165–168) Type material. Holotype: CHINA (YUNNAN Prov.): ♂, two labels: “CHINA mer.: Yunnan prov. / (pass SW from Baoshan / Gaoligong Shan) / 4. - 8.6.2005 / sifted from base of old tree / Oto Nakladal lgt.” [white, printed]; “ CEPHENNODES (s. str.) / dilatatus m. / P. Jałoszyński, 2025 / HOLOTYPUS ” [red, printed] (MNHW). Diagnosis. Body small, BL around 1.2 mm; antennae gradually thickened; punctures on clypeus and frontovertexal region very small but sharply marked and dense, separated by spaces subequal to their diameters, pronotum with slightly larger, more distinct and similarly dense punctures, elytra with punctures similar to those on head; subhumeral lines as long as about 0.2 × EL. Male: abdomen (Fig. 166) with sternite V modified, bearing conspicuously broad median subtriangular transverse carina; metatibia (Fig. 165) strongly thickened in subapical region; aedeagus (Figs 167–168) in ventral view broadest in subapical region, apical projection oblique in relation to long axis of median lobe, its apex not hidden behind apex of capsular region of median lobe but adpressed against it. Description. Body of male (Fig. 165) strongly convex, oval, with indistinct constriction between pronotum and elytra, moderately dark brown with lighter tarsi, covered with light brown setae. BL 1.20 mm. Head broadest across moderately large and moderately strongly convex, coarsely faceted eyes, HL 0.18 mm, HW 0.28 mm; frontovertexal region weakly convex at middle; supraantennal tubercles weakly elevated. Punctures on clypeus and frontovertexal region very small but sharply marked and dense, separated by spaces subequal to their diameters; setae sparse, short, suberect. Antennae moderately long and slender, gradually thickened, AnL 0.60 mm, antennomere 1 about as long as broad, 2 strongly elongate, 3 distinctly transverse, 4–7 each about as long as broad, 8–10 each weakly transverse, 11 as long as 9 and 10 combined, about 1.8 × as long as broad. Pronotum semi-oval, strongly convex at middle and flattened near hind corners, broadest near middle and distinctly narrowing posterad; PL 0.35 mm, PW 0.50 mm. Anterior margin in strictly dorsal view nearly straight; lateral margins strongly rounded in anterior half and less strongly posteriorly, finely microserrate, posterior corners nearly right-angled; posterior margin shallowly bisinuate; lateral marginal carinae simple (i.e., not doubled); lateral antebasal pits shallow but distinct, each slightly closer to lateral than posterior pronotal margin. Punctures on pronotal disc slightly larger and deeper than those on head, distinct, separated by spaces subequal to their diameters; setae short and moderately dense, suberect. Elytra slightly wider than pronotum, together oval, broadest near anterior third; EL 0.68 mm, EW 0.55 mm, EI 1.23; subhumeral lines as long as about 0.2 × EL, developed as sharp borders between elevated humerus and lower mesal region, strongly divergent posterad; basal elytral fovea on each elytron situated slightly closer to lateral margin of mesoscutellum than to subhumeral line; elytral apices separately rounded. Punctures on elytra similar to those on head, shallower and slightly sparser than those on pronotum; setae similar to those on pronotum. Hind wings not studied. Metaventrite with lateral impressions as long as about half of its length. Legs long and slender, metatibiae with wide subtriangular outer lateral subapical expansion (Fig. 165). Abdomen (Fig. 166) with sternite V bearing long transverse subtriangular median carina. Aedeagus (Figs 167–168) of simonis form, AeL 0.14 mm; median lobe in ventral view broadest in subapical region, apical projection oblique in relation to long axis of median lobe, its apex not hidden behind apex of capsular region of median lobe but adpressed against it; parameres long and slender, one nearly reaching apex of aedeagus, each with one short apical seta. Female. Unknown. Distribution. S China: Yunnan. Etymology. The adjective dilatatus refers to the dilated hind tibiae. Remarks. Among Chinese species, C. dilatatus can be identified by a combination of the following characters: aedeagus in ventral view broadest in subapical region, apical projection oblique in relation to long axis of median lobe, its apex not hidden behind apex of capsular region of median lobe but adpressed against it. Among the Himalayan species, C. dolakhanus, C. manangensis, C. monolaminatus, and C. thakanus have a modified sternite V and unmodified IV, as in C. dilatatus. Cephennodes dolakhanus and C. thakanus have clearly different shapes of the apical plates of the median lobe. Cephennodes manangensis and C. monolaminatus have the aedeagi most similar to that of C. dilatatus. Cephennodes manangensis has a different pattern of punctures on body parts: on the pronotal disc punctures are much smaller and shallower than those on the head, but denser, and on elytra slightly more distinct than those on pronotum, whereas in C. dilatatus the pronotum is covered with punctures slightly larger, more distinct and similarly dense as those on the frontovertexal region, and elytra with punctures similar to those on the head. Cephennodes monolaminatus is distinctly larger than C. dilatatus, body length respectively 1.45–1.58 mm and 1.20 mm, and has the pronotal disc covered with fine and inconspicuous punctures.

Zenodo (CERN European Organization for Nuclear Research)
Life below water
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Subterranean biodiversity and taxonomy
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