Justicia werffii J. R. I. Wood & R. Villanueva 2025, sp. nov.

34.Justicia werffiiJ. R. I. Wood & R. Villanueva sp. nov. Type. Peru • San Martin, Prov. Rioja, Carretera Rioja [6°03'S, 77°10'W] – Pedro Ruiz [5°57'S, 77°58'W], 1450 m, 2 March 1998, H. Van der Werff 15565 (holotype MO-5763942, isotypes US-3387908, USM). Diagnosis. A small-flowered species that resembles J. morona-santiagoensis Wassh. in the spathulate bracts but differs in the liana habit (not erect or ascending subshrub), short peduncles, 0.5–2 cm long (not 4–6 cm) and the ciliate bracts and bracteoles (not glabrous). Description. Liana of unknown height; stems sulcate, glabrous. Leaves petiolate, lamina 2.5–6 × 1–2.3 cm, apex acuminate, base cuneate, slightly oblique, margin entire to crenulate, subglabrous or with few hairs in the veins, numerous small cystoliths present, veins 5–6 pairs; petioles 3–7 mm, subglabrous or hirtellous. Inflorescence of solitary or paired axillary racemes 1–4 cm long, becoming aggregated apically to form a terminal thyrse, the leaves progressively reduced in size; peduncles 0.5–2 cm, glabrous; rhachis 0.5–2 cm long, scurfy; flowers on pedicels 0–2 mm long; bracts and bracteoles similar, spathulate, 4 × 1.5 mm, ciliolate; calyx 5 - lobed to c. 1 mm above base, lobes 4 × 0.5 mm, lanceolate, acuminate, puberulent; corolla c. 12 mm long, white, minutely tomentellous, the tube c. 3.5 mm, the base somewhat bulbous, upper lip c. 8 mm long, minutely notched, erect, lower lip c. 6 mm, 3 - lobed, lobes c. 2–2.5 × 1.5–2 mm, narrowly ovate, obtuse; anthers shortly exserted, thecae ellipsoid, c. 1.5 × 1 mm, slightly superposed, glabrous, basally acute; pollen prolate, 29–35 × 16–21 μm, 3 - aperturate, colporate, 1 row of ± 7–8 distinct insulae on either side of aperture (Fig. 51 B); style 12 mm, glabrous except for a few hairs at base; ovary glabrous. Capsule and seeds not seen. Illustration. Fig. 33. Etymology. This species is named for Henk van der Werff, who collected the type as well as many other important specimens from northern Peru. Phenology. Found in flower in March. Habitat. Montane forest on clay soil in San Martin, 1450 m. Distribution. Endemic to Peru and only known from the type, which was collected in San Martin. Fig. 64. Material examined. Peru • San Martin: type collection.