Talopena apicina, comb. nov.

Talopena apicina (Gould, 1861) comb. nov. Figs 35–38 Monilea apicina Gould, 1861: 16. Type locality: Port Jackson [Sydney Harbour, New South Wales], leg. W. Stimpson; erroneous – here emended to Lansdowne Bank, eastern Coral Sea [20.167°S, 160.617°E]. Monilea apicina – Gould 1862: 154. — Pilsbry 1889–1890: 254. — Iredale 1924: 228. — Johnson 1964: 41, pl. 14 fig. 4 (holotype) Isanda (Parminolia) apicina – Ladd 1966 (partim): 39, pl. 5 figs 3–4 (holotype). Monilea incerta (non Iredale, 1912) – Héros et al. 2007: 209. Parminolia apicina – Wilson 1993: 94. Non Parminolia apicina – Iredale 1929a: 271. — Héros et al. 2007: 209 [= Parminolia agapeta (Melvill & Standen, 1896)]. Material examined Holotype (Fig. 38A–D) AUSTRALIA • Port Jackson [erroneous – see Remarks] USNM 24159. Paratype (Fig. 38E) AUSTRALIA • Port Jackson [erroneous] NHMUK 20210269. Other material NEW CALEDONIA – Plateau Chesterfield-Bellona • 4 specimens, living; Plateau des Chesterfield, Stn DW147; 19°37′ S, 158°14′ E; depth 25 m; 28 Jul. 1988; NO Coriolis -CORAIL 2 leg.; MNHN • 2 specimens, dead; Plateau des Chesterfield, Stn DW136; 19°31′ S, 158°16′ E; depth 37 m; 30 Jul. 1988; NO Coriolis -CORAIL 2 leg.; MNHN • 2 specimens, dead; Plateau des Chesterfield, Stn DW48; 19°18′ S, 158°27′ E; depth 44 m; 23 Jul. 1988; NO Coriolis -CORAIL 2 leg.; MNHN • 2 specimens, dead; Plateau des Chesterfield, Stn DW42; 19°22′ S, 158°29′ E; depth 45 m; 23 Jul. 1988; NO Coriolis -CORAIL 2 leg.; MNHN • 14 specimens, dead; Plateau des Chesterfield, Stn DW128; 19°28′ S, 158°30′ E; depth 38 m; 29 Jul. 1988; NO Coriolis -CORAIL 2 leg.; MNHN • 4 specimens, living; Plateau Chesterfield-Bellona, Stn D45; 20°48.93′ S, 158°30.21′ E; depth 50 m; Jul. 1984; CHALCAL 1 leg.; MNHN • 1 specimen, dead; Plateau des Chesterfield, Stn DW76; 19°12′ S, 158°33′ E; depth 53 m; 25 Jul. 1988; NO Coriolis -CORAIL 2 leg.; MNHN • 1 specimen, dead; Plateau Chesterfield-Bellona, Stn D26; 19°10.72′ S, 158°34.95′ E; depth 48 m; Jul. 1984; CHALCAL 1 leg.; MNHN • 1 specimen, dead; Plateau Chesterfield-Bellona, Stn D49; 20°58.2′ S, 158°35.0′ E; depth 48 m; Jul. 1984; CHALCAL 1 leg.; MNHN • 10 specimens, dead; Plateau des Chesterfield, Stn D39; 20°28.9′ S, 158°40.7′ E; depth 40 m; Jul. 1984; CHALCAL 1 leg.; MNHN • 1 specimen, dead; Plateau des Chesterfield, Stn DW38; 19°22′ S, 158°43′ E; depth 61 m; 23 Jul. 1988; NO Coriolis -CORAIL 2 leg.; MNHN • 1 specimen, dead; Plateau des Chesterfield, Stn DW79; 19°12′ S, 158°45′ E; depth 58 m; 25 Jul. 1988; NO Coriolis -CORAIL 2 leg.; MNHN • 1 specimen, dead; Plateau des Chesterfield, Stn DW31; 19°25′ S, 158°45′ E; depth 57 m; 23 Jul. 1988; NO Coriolis -CORAIL 2 leg.; MNHN • 1 specimen, dead; Plateau Chesterfield-Bellona, Stn D37; 19°54′ S, 158°46.3′ E; depth 50 m; Jul. 1984; CHALCAL 1 leg.; MNHN • 23 specimens, dead; Plateau Chesterfield-Bellona, Stn D17; 19°11.9′ S, 158°55.8′ E; depth 44 m; Jul. 1984; CHALCAL 1 leg.; MNHN • 11 specimens, living; Plateau des Chesterfield, Stn DW110; 19°09′ S, 158°56′ E; depth 40 m; 28 Jul. 1988; NO Coriolis -CORAIL 2 leg.; MNHN • 4 specimens, dead; Plateau Chesterfield-Bellona, Stn DW85; 19°12′ S, 158°56′ E; depth 32 m; 26 Jul. 1984; CHALCAL 1 leg.; MNHN • 4 specimens, dead; without further detail; CHALCAL 1 leg.; MNHN. – Lansdowne-Fairway Banks • 2 specimens, dead; Bancs Lansdowne–Fairway, Stn D8; 20°47.3′ S, 161°01.4′ E; depth 40 m; Jul. 1984; CHALCAL 1 leg.; MNHN • 1 specimen, dead; Bancs Lansdowne–Fairway, Stn DW26; 20°22′ S, 161°05′ E; depth 62 m; 22 Jul. 1988; NO Coriolis -CORAIL 2 leg.; MNHN • 2 specimens, dead; Bancs Lansdowne– Fairway, Stn DW02; 20°50′ S, 161°37′ E; depth 62 m; 20 Jul. 1988; NO Coriolis -CORAIL 2 leg.; MNHN. – d’Entrecasteaux Reefs – Lagon Nord – Belep • 1 specimen, dead; Atoll de Huon, Stn 436; 18°06′ S, 162°50′ E; depth 45 m; 25 Feb. 1985; B. Richer-ORSTOM leg.; MNHN • 1 specimen, dead; Lagon Nord, Stn 488; 18°53′ S, 163°30′ E; depth 38 m; 2 Mar. 1985; B. Richer-ORSTOM leg.; MNHN • 1 specimen, dead; Lagon Nord, Stn 487; 18°55′ S, 163°31′ E; depth 37 m; 2 Mar. 1985; B. Richer-ORSTOM leg.; MNHN • 3 specimens, dead; Lagon Nord, Stn 481; 18°57′ S, 163°32′ E; depth 33 m; 2 Mar. 1985; B. Richer-ORSTOM leg.; MNHN • 1 specimen, dead; Lagon Nord, Stn 518; 19°05′ S, 163°35′ E; depth 38 m; 5 Mar. 1985; B. Richer-ORSTOM leg.; MNHN • 2 specimens, living; Stn 1105; 19°40′ S, 163°57′ E; depth 25 m; 25 Oct. 1989; B. Richer-ORSTOM leg.; MNHN • 12 specimens, living; Stn 1104; 19°42′ S, 163°59′ E; depth 22 m; 25 Oct. 1989; B. Richer-ORSTOM leg.; MNHN. – Grande Terre, Koumac • 1 specimen, dead; Stn 947; 20°33′ S, 164°07′ E; depth 17–18 m; 28 Apr. 1988; B. Richer-ORSTOM leg.; MNHN • 1 specimen, living; Chenal de l’Infernet, Stn 1305; 20°36.2′ S, 164°11.0′ E; depth 12–15 m; Oct. 1993; Expédition Montrouzier leg.; sable gris; MNHN • 1 specimen, dead; Chenal de l’Infernet, Stn 1306; 20°39.1′ S, 164°12.4′ E; depth 11–13 m; Oct. 1993; Expédition Montrouzier leg.; fonds blancs; MNHN • 5 specimens, living; Chenal de l’Infernet, Stn 1304; 20°38.6′ S, 164°13.2′ E; depth 12–15 m; Oct. 1993; Expédition Montrouzier leg.; sable gris, dalle; MNHN • 1 specimen, dead; Stn KD568; 20°40.1′ S, 164°14.3′ E; depth 2 m; 16 Nov. 2019; KOUMAC 2.3 leg.; sable blanc et quelques débris coralliens; MNHN • 2 specimens, dead; Stn KD567; 20°40.1′ S, 164°14.6′ E; depth 2 m; 16 Nov. 2019; KOUMAC 2.3 leg.; sable blanc et algues vertes filamenteuses; MNHN • 1 specimen, dead; Chenal de la Passe de Koumac, Stn 1315; 20°40.7′ S, 164°14.7′ E; depth 66–87 m; Oct. 1993; Expédition Montrouzier leg.; sable coquillier vaseux; MNHN • 8 specimens, dead; Chenal de la Passe de Koumac, Stn 1323; 20°40.9′ S, 164°14. 8′ E; depth 82–120 m; Oct. 1993; Expédition Montrouzier leg.; sable coquillier vaseux; MNHN • 7 specimens, living; Stn KD560; 20°45.6′ S, 164°15.2′ E; depth 5–9 m; 14 Nov. 2019; KOUMAC 2.3 leg.; sable blanc et quelques algues brunes; MNHN • 2 specimens, living; Stn KD559; 20°46′ S, 164°15.5′ E; depth 12–14 m; 14 Nov. 2019; KOUMAC 2.3 leg.; sable blanc; DNA voucher; MNHN-IM-2019-8826 • 1 specimen, living; Stn KD572; 20°45.8′ S, 164°16.1′ E; depth 22–23 m; 18 Nov. 2019; KOUMAC 2.3 leg.; sable grossier, coquilles; MNHN • 2 specimens, living; Stn KD554; 20°47′ S, 164°16.8′ E; depth 2–5 m; 13 Nov. 2019; KOUMAC 2.3 leg.; sable blanc et gros débris de coraux morts; MNHN • 2 specimens, living; Stn KD553; 20°47.9′ S, 164°17.2′ E; depth 5–10 m; 13 Nov. 2019; KOUMAC 2.3 leg.: sable blanc; DNA voucher; MNHN-IM- 2019-8778. – Grande Terre, Bourail • 7 specimens, dead; Lagon de Poé, Stn 1324; 21°36.9′ S, 165°22.7′ E; depth 0–2 m; 1 Sep. 1993; Expédition Montrouzier leg.; fonds mixtes, herbier; MNHN. – Grande Terre, Baie de St Vincent • 1 specimen, dead; Stn 186; 22°04′ S, 166°00′ E; depth 11 m; 19 Sep. 1984; B. Richer-ORSTOM leg.; MNHN • 4 specimens, dead; Stn 162; 22°13′ S, 166°09′ E; depth 10 m; 18 Sep. 1984; B. Richer-ORSTOM leg.; MNHN. – Grande Terre, Nouméa • 9 specimens, living; Stn DE56; 22°08.4′ S, 166°07.7′ E; depth 19–20 m; 29 Apr. 1987; B. Richer de Forges-IRD leg.; MNHN • 3 specimens, dead; Stn 51; 22°15′ S, 166°11′ E; depth 10 m; 25 May 1984; B. Richer-ORSTOM leg.; MNHN • 1 specimen, dead; Stn 50; 22°17′ S, 166°12′ E; depth 12 m; 25 May 1984; B. RicherORSTOM leg.; MNHN • 1 specimen, dead; Stn 49; 22°19′ S, 166°14′ E; depth 10 m; 25 May 1984; B. Richer-ORSTOM leg.; MNHN • 11 specimens, living; Grand Récif Aboré, Stn 1346; 22°21.9′ S, 166°16.1′ E; depth 5–6 m; 5 Oct. 1992; P. Bouchet leg.; fonds blanc; MNHN • 1 specimen, living; Stn DE62; 22°16.8′ S, 166°16.6′ E; depth 32–34 m; 30 Apr. 1987; B. Richer de Forges-IRD leg.; MNHN • 5 specimens, living; Stn 8; 22°23′ S, 166°18′ E; depth 15 m; 21 May 1984; B. Richer-ORSTOM leg.; MNHN • 1 specimen, dead; Quatre Bancs de l’Ouest, Stn 1511; 22°26.5′ S, 166°18.3′ E; depth 0–1 m; 26 Oct. 2000; Cosel and Trondlé leg.; platier, marée basse, récolte à vue; MNHN • 7 specimens, living; Quatre Bancs de l’Ouest, Stn 1510; 22°26.47′ S, 166°18.35′ E; depth 0 m; 26 Oct. 2000; Cosel and Trondlé leg.; hand dredge; MNHN • 1 specimen, dead; Stn 257; 22°22′ S, 166°20′ E; depth 9 m; 7 Nov. 1984; B. Richer-ORSTOM leg.; MNHN • 1 specimen, dead; Stn 7; 22°24′ S, 166°20′ E; depth 14 m; 21 May 1984; B. Richer-ORSTOM leg.; MNHN • 10 specimens, dead; Grand Récif Aboré, Stn 1347; 22°23.6′ S, 166°20.1′ E; depth 10 m; 8 Oct. 1992; P. Bouchet leg.; vase sableuse sur dalle; MNHN • 5 specimens, dead; Île aux Goélands, Stn 1368; 22°24.3′ S, 166°20.7′ E; depth 10 m; 16 Apr. 1993; P. Bouchet leg.; fonds blancs; MNHN • 4 specimens, living; Stn 4; 22°23′ S, 166°21′ E; depth 9 m; 21 May 1984; B. Richer-ORSTOM leg.; MNHN • 40 specimens, living; Crouy; 22°21.5′ S, 166°21.7′ E; depth 1–6 m; 1990–2003; Claude Berthault leg.; MNHN • 7 specimens, living; SE Sèche Croissant, Stn 1508; 22°18.6′ S, 166°21.8′ E; depth unknown; 25 Oct. 2000; Cosel and Trondlé leg.; hand dredged; MNHN • 10 specimens, dead; Île aux Goélands, Stn 1503; 22°22.4′ S, 166°22.5′ E; depth 5–7 m; 21 Oct. 2000; Cosel and Trondlé leg.; MNHN • 1 specimen, living; Stn 21; 22°23′ S, 166°23′ E; depth 10 m; 23 May 1984; B. Richer-ORSTOM leg.; MNHN • 1 specimen, living; Stn 22; 22°26′ S, 166°23′ E; depth 11 m; 23 May 1984; B. Richer-ORSTOM leg.; MNHN • 2 specimens, dead; Stn 281; 22°24′ S, 166°24′ E; depth 10 m; 9 Nov. 1984; B. Richer-ORSTOM leg.; MNHN • 1 specimen, dead; Stn 283; 22°27′ S, 166°24′ E; depth 13 m; 9 Nov. 1984; B. Richer-ORSTOM leg.; MNHN • 1 specimen, dead; Ile Ouen-Baie du Prony, Stn 64; 22°28′ S, 166°25′ E; depth 15 m; 20 Aug. 1984; B. Richer-ORSTOM leg.; MNHN • 2 specimens, living; Anse Vata; 22°18.1′ S, 166°26.4′ E; depth 1–5 m; 1987–2007; Claude Berthault leg.; MNHN • 2 specimens, dead; Stn DE38; 22°33.6′ S, 166°28.6′ E; depth 22–24 m; 29 Apr. 1987; B. Richer de Forges-IRD leg.; MNHN • 1 specimen, dead; Ile Ouen-Baie du Prony, Stn 82; 22°33′ S, 166°29′ E; depth 10 m; 21 Aug. 1984; B. Richer-ORSTOM leg.; MNHN • 6 specimens, living; Stn DE36; 22°35.5′ S, 166°32.0′ E; depth 17–19 m; 2