Coleus scruposus A. J. Paton

Coleus scruposus A. J. Paton, Phytokeys 129: 92. 2019. ≡ Pycnostachys kassneri De Wild., Contr. Fl. Katanga: 172. & Ann. Soc. Sci. Bruxelles 41 (2): 54. 1921., non Coleus kassneri Robyns & Lebrun. = Pycnostachys congensis Gürke, Bull. Herb. Boissier 4: 819. 1896. Type: DR. Congo, Lusambo, 1891, G. Descamps 27 (lectotype Z [Z- 000021015], designated here). ≡ Coleus congensis (Gürke) A. J. Paton, Phytokeys 129: 35. 2019. Type. DR. Congo, Katanga, West Kundelungu, 17 May 1908, T. Kassner 2794 (holotype BR [BR 0000009824614]; isotype BM [BM 000838018], HBG, K [K 000405735]). Description. Paton et al. (2009: 390), Paton et al. (2013: 318), as Pycnostachys kassneri De Wild. Distribution. Tanzania to Zambia. Habitat and еcology. Savannah, riparian forest, scree, dry woodland; 950–1650 m elev. Additional specimens. DR. Congo, Kasaï, Sandoa, près de Kambungu, 23 Apr 1959, S. Risopoulos 970 (BR, K, WAG); Bas-Katanga, Samba, 1891, G. Descamps 28 (BR); Kamina, Lovoi, Mar 1932, P. Quarré 2971 (BR); Haut-Katanga, Upemba, Mont Muye, 3 May 1948, G. F. de Witte 3774 (BR); Vallée de la Kalumengongo, près des chutes, 12 Apr 1949, G. F. de Witte 6071 (BR, K, WAG); Mugila, 23 May 1908, T. Kassner 2967 (K); Kundelungu, 17 km NW Sampwe, 24 Apr 1970, S. Lisowski, F. Malaisse, J. - J. Symoens 11072 (BR, POZG); Route Kolwezi-Kamina, km 75, 5 Apr 1948, A. Schmitz 1780 (BR). Notes. 1. Calyx teeth length is variable; specimens from DR. Congo and Zambia with very short calyx teeth (<2 mm) also tend to have calyx throat that is tomentose inside (e. g. A. Schmitz 1780, J. - J. Symoens 10239). Further work is needed to decide if this represents a distinct taxon or just one extreme of a continuous variation. 2. Lectotypification of Pycnostachys congensis Gürke. Gürke (1896) cited two syntypes of Pycnostachys congensis Gürke (G. Descamps 27 (syntype B destroyed; isosyntype Z), DR. Congo, Lusambo & G. Descamps 28 (syntype B destroyed; isotype BR), DR. Congo, Samba, 1891. We select G. Descamps 27 (Z) as the lectotype because it comprises a whole plant.