Polana elera DeLong & Freytag 1972, comb. nov.

Polana elera DeLong & Freytag, 1972 comb. nov. Polana (Largulara) elera DeLong & Freytag, 1972: 293. Polana (Varpulana) elera —Freytag, 1992: 140. Remarks. Similar to P. ardua, the male genitalia of Polana elera (DeLong & Freytag, 1972: 294, figs 296–298) is identical to the genitalia of Effossana tenuemarginata Domahovski & Cavichioli, 2023 (see figures 41G–M in Domahovski & Cavichioli, 2023: 146), but with distinctly smaller length of body and different coloration. In addition, its male genitalia strongly contrast with the other species of Varpulana as noted by Domahovski & Cavichioli (2017). Therefore, this species is here considered as incertae sedis within Polana. DeLong & Triplehorn (1979) described five species of Polana from Peru, placing four in the subgenus Nihilana. However, two of these species have a pair of processes arising from the basal portion of the aedeagus and therefore cannot be placed in Nihilana, because one of the diagnostic features of Nihilana is the absence of the basal pair of processes (called ‘parameres’ by the authors). Both subgenera, Nihilana and Polana have the pygofer with basodorsal hooked-shape processes (DeLong & Freytag 1972). The new combinations are as follows: