Odorrana lipuensis Mo, Chen, Wu, Zhang & Zhou 2015

Odorrana lipuensis Mo, Chen, Wu, Zhang & Zhou, 2015 Related literature. Mo et al. (2015); Pham et al. (2016 a, 2017); Poyarkov et al. (2021); Luong et al. (2022). Type materials. Holotype. NHMG 1306001, adult male, from a completely dark karst cave (24°38'N, 110°26'E; 182 m a. s. l.) in Maling Town, Lipu City, Guangxi Zhuang Autonomous Region, China. Paratypes. (n = 7): NHMG 1306002, NHMG 20140702, NHMG 20140703, adult males and NHMG 1303018, NHMG 1303019, NHMG 1306003, NHMG 20140701, adult females, from the same locality as the holotype. Additional specimen. SYS a 004898, adult female, from Yangshuo County, Guangxi Zhuang Autonomous Region, China. Variation. Our morphological analyses of the additional specimen of this species provided new morphometric data (Table 2) and suggested the overall morphology of this specimen is generally consistent with previous descriptions (Mo et al. 2015; Pham et al. 2016 a; Pham et al. 2017). However, a morphological revision was also suggested. Our morphological examination revealed fingers II – IV and toes I – V with lateroventral grooves in the examined specimen, which differ from the previous description of all fingers and toes with circum-marginal grooves (Mo et al. 2015; Pham et al. 2016 a; Pham et al. 2017). Revised diagnosis. This species can be diagnosed from other species of Odorrana lipuensis group by the combination of following morphological characteristics: (1) body size moderate, SVL 40.7–49.8 mm in adult males (n = 6) and 49.9–60.1 mm in adult females (n = 8); (2) relative finger length I ≥ II; (3) dorsolateral conical spines dense; (4) dorsolateral folds absent in both sexes; (5) bands on the dorsal surface of hind-limb clear; and (6) nuptial pad present on finger I. Distribution. This species is known from several localities in China and Vietnam, including: (1) Maling Town, Lipu City, Guangxi, (2) Yangshuo Town, Yangshuo County, Guangxi and (3) Ha Lang District, Cao Bang (Fig. 1). The record of this species in Bac Kan, Vietnam was tentatively unconfirmed (Fig. 1) for the reason outlined in Remarks under Odorrana liboensis.