Enica longirostris

Enica longirostris (Wiedemann, 1819) Fig. 10 Anthrax longirostris Wiedemann, 1819: 11. Cyllenia afra Wiedemann, 1828: 358. Cyllenia pluricellata Macquart, 1855: 104. Taxon depository. ZooBank: https://zoobank.org/b4e3556d-a8d8-4dd0-924b-7b7f8689dab2. Diagnosis. The species is distinguished from other species in the genus by the two crossveins connecting R 1 and R 2 + 3 and by densely setose gena. Description. Head: wider than high, creamy white; males holoptic, females dichoptic, vertex and compound eyes at same level; frons (at level of antennal insertion) more or less parallel-sided, female: light brown to reddish-orange, darker than head, without dark spot, males: similar to head, slightly darker than head; frons black setose, with white scales, broadly setose near eyes, extending ventral to antennae; ocellar tubercle slightly raised, dark reddish brown to black, black macrosetose, white or light brown scales posteriorly; occiput whitish – yellow setose, brown setose on occiput spots, white scales antero-dorsally, antero-dorsally with dark brown to black spots, usually oval; gena creamy white, densely white setose. Proboscis and maxillary palpus: proboscis straight, black; labella only forming distal tip of proboscis, apically rounded, minute black setose; maxillary palpus not extending beyond oral cavity, light brown to brown, brown setose or light yellowish-brown setose. Antenna: dark brown; scape creamy white, approximately as long as pedicel, variable setation color: black setose, white setose, or white scales may be present, setae extending to tip of pedicel; pedicel reddish brown or brown, short black setose dorsally; postpedicel black or dark brown, tapering distally, longer than scape and pedicel combined, white or light yellow scales dorsally; stylus reduced, only apical ‘ seta-like’ sensory element present, situated sub – apically in cavity on postpedicel. Thorax: light orangish brown or reddish brown and light orangish brown; scutum predominantly black, reddish on margins, light to dark brown scales, two submedial longitudinal stripes with lighter scales; setation: acr setae present, black, dc setae present, black; prosternum silvery grey pubescence; proepisternum silvery grey pubescence, long yellowish-white setose, white scales; antepronotum silvery grey pubescence, dull white scales anteriorly; postpronotum silvery grey pubescence, asetose, with medial dark brown to black stripe; postpronotal lobe silvery grey pubescence, white scales, white and black setose; pleuron silvery grey pubescence; proepimeron silvery grey pubescence, asetose; anepisternum silvery grey pubescence, white scales ventrally, black macrosetose dorsally; anepimeron silvery grey pubescence, asetose or white scales; katepisternum silvery grey pubescence, white scales dorsally; laterotergite silvery grey pubescence, asetose; meron + metepisternum silvery grey pubescence, asetose or white scales dorsally; metepimeron silvery grey pubescence, asetose or white setose dorsally; scutellum red, sometimes with anterior or posterior black spots, apubescent, ds sctl setae black macrosetose, ap sctl setae comprised of light brown and white scales. Leg: light orangish brown, covered in scales, some regular setae present; pro coxa light orangish brown, silvery grey pubescence, white scales, thick black setose; pro femur light orangish brown, white scales ventrally, brown scales dorsally, few short black setae; pro tibia light brown, raised short and fine dark brown setose ventrally, large black setose dorsally; mes coxa light orangish brown, white pubescence, white scales, thick black setose; mes femur light orangish brown, white scales ventrally, brown scales dorsally, few short black setae; mes tibia light brown, dark brown scales, long black setose, distal tip with long black setae; met coxa light orangish brown, white pubescent, white scales, thick black setose, anteriorly without any protuberance; met trochanter short black setose ventrally, white scales dorsally, without protuberance; met femur light orangish brown, white scales ventrally, brown scales dorsally, few short black setae; met tibia light brown, dark brown scales, large black setose, distal tip with long black macrosetae of varying sizes; proximal pro, mes, and met tarsomeres longer than following 2 tarsomeres combined, proximal met tarsomere as wide as following tarsomeres; pro tarsomere with dark brown scales dorsally, black setose ventrally; mes tarsomere with dark brown scales dorsally, black setose ventrally; met tarsomere with dark brown scales dorsally, black setose ventrally; claw smoothly arched distally. Wing: (6.1 –) 9.1–9.8 (– 11.4) mm, microtrichia absent; membrane color pattern males: predominantly brown colored except for white tinted base of wing, supernumerary crossveins between R 1 and R 2 + 3, oblique rectangular area between CuA and R 1, crossveins r-r and r-m, and base of cell m 2, females: membrane dark brown colored except for white tinted base of wing, supernumerary crossveins between R 1 and R 2 + 3, oblique rectangular area between CuA and R 1, crossveins r-r and r-m, and base of cell m 2; costal vein dark brown short setose, whitish clear setose proximally; R 2 + 3 distally distinctly arching anteriorly, r 1 open, divided by 2 supernumerary crossveins; R 4 terminating anterior to wing apex, distinctly arching anteriorly; cell r 4 open, proximally R 4 and R 5 parallel, R 4 strongly diverging anteriorly in distal 2 / 3; R 5 terminating posterior to wing apex; r 5 open; M 1 terminating posterior to wing apex; cell d closed by base of M 2 and m-m, r-m situated in distal ¼; m 3 open; cua open; alula greatly reduced, nearly straight wing margin; haltere stem dark brown setose. Abdomen: shape ovate, black, predominantly black anteriorly and red posteriorly or reddish brown; tergites smooth, setae with small sockets only; T 1 reddish yellow to light brown, reddish brown, or predominantly black, no red or a very thin line of red posteriorly, light yellowish-brown to light whitish brown setose, scales or snow – white setose, with scales; T 2 – T 7 predominantly reddish-orange or predominantly black or predominantly dark reddish brown, light brown setose, white scales, rarely brown to light yellowish-brown setose, white scales or dark brown setose, white scales, white scales on medial spots on T 2–8 and posterior margins of T 2 and T 4, rarely scales uniformly color throughout or light brown setose, 2 spots with dark scales on T 2 and T 3; S 1–8 predominantly black with reddish brown, yellow to white scales, thin black setose and long white setose or brown scales, thin black setose and long white setose; lateral margin lacking setae, rarely sparse black setae on T 4–8, male T 8 white scales, black setose. Terminalia: Male terminalia drawings in Hesse (1956, p 32): https://www.biodiversitylibrary.org/page/40844167. Other material. South Africa – Eastern Cape • 1 ♀ Alicedale, 2 km E; 33°19'07"S, 026°06'12"E; 310 m a. s. l.; 23 Oct. 2004; Londt, Jason leg.; Dry rocky hillside with succulent vegetation; NMSA -DIP-93654, NMSA • 1 ♂ Alicedale, 2 km E; 33°19'07"S, 026°06'12"E; 310 m a. s. l.; 23 Oct. 2004; Londt, Jason leg.; Dry rocky hillside with succulent vegetation; NMSA -DIP-93655, NMSA • 1 ♀ Alicedale, 2 km E; 33°19'07"S, 026°06'12"E; 310 m a. s. l.; 23 Oct. 2004; Londt, Jason leg.; Dry rocky hillside with succulent vegetation; NMSA -DIP-93656, NMSA • 1 ♀ 3 ♂ 1? Groendal, Uitenhage (= kariega); 33°43'13"S, 025°18'55"E; 28 Oct. 1938; SAM Museum Staff leg.; SAM-DIP-A 002141, SAMC • 1 ♀ Willowmore; 33°17'00"S, 023°29'00"E; 25 Oct. 1918; Brauns, H. leg.; NMSA -DIP-054408, NMSA • 1 ♀ Willowmore; 33°17'00"S, 023°29'00"E; 01 Nov. 1919; Brauns, H. leg.; NMSA -DIP-054687, NMSA • 1 ♀ Willowmore; 33°17'00"S, 023°29'00"E; 01 Nov. 1919; Brauns, H. leg.; NMSA -DIP-89485, NMSA • 1 ♂ Willowmore; 33°17'00"S, 023°29'00"E; 15 Oct. 1921; Brauns, H. leg.; NMSA -DIP-056255, NMSA • 1 ♂ Willowmore; 33°17'00"S, 023°29'00"E; 15 Oct. 1921; Brauns, H. leg.; NMSA -DIP-89530, NMSA • 1 ♀ Willowmore; 33°17'00"S, 023°29'00"E; Dec. 1922; Brauns, H. leg.; NMSA -DIP-89522, NMSA • 1 ♂ Willowmore; 33°17'00"S, 023°29'00"E; Dec. 1922; Brauns, H. leg.; NMSA -DIP-89523, NMSA South Africa – Northern Cape • 1 ♀ Bowesdorp (= Bowes Dorp), Namaqualand; 30°09'03"S, 017°55'55"E; Nov. 1931; SAM Museum Staff leg.; SAM-DIP-A 002167, SAMC • 2 ♀ Hester Malan N. R. (= Goegap Nature Reserve), Namaqualand; 29°40'00"S, 018°02'00"E; 16 Oct. 1985; Struck, M. leg.; SAM-DIP-A 004968, SAMC • 1 ♀ Kamieskroon Namaqualaud; 30°12'00"S, 017°56'00"E; Nov. 1936; SAM Museum Staff leg.; SAM-DIP-A 002143, SAMC • 1 ♀ Kamieskroon, 10 km E; 30°11'39"S, 018°00'10"E; 630 m a. s. l.; 17 Oct. 1977; Miller, R. M. leg.; NMSA -DIP-049597, NMSA • 1 ♂ Nieuwoudtville; 31°23'00"S, 019°06'00"E; Aug. 1928; Brauns, H. leg.; NMSA -DIP-056002, NMSA • 1 ♂ Nieuwoudtville, 1 km W, caravan park and area; 31°23'00"S, 019°06'00"E; 800 m a. s. l.; 04 Nov. 1991; Londt, Jason leg.; NMSA -DIP-92784, NMSA • 1 ♀ Nieuwoudtville, 1 km W, caravan park and area; 31°23'00"S, 019°06'00"E; 800 m a. s. l.; 04 Nov. 1991; Londt, Jason leg.; NMSA -DIP-92785, NMSA • 1 ♀ Nieuwoudtville, 9 km W, Summit Vanrhyn’s Pass; 31°22'00"S, 019°01'00"E; 700 m a. s. l.; 04 Nov. 1991; Londt, Jason leg.; NMSA -DIP-92829, NMSA • 1 ♂ Outiep Garies Namaqualand (= Namakwaland); 30°33'27"S, 017°59'34"E; Sep. 1953; Toit, J. leg.; SAM-DIP-A 002145, SAMC • 1 ♀ Springbok, 8 km NW, road to Nababeep; 29°37'16"S, 017°49'59"E; 985 m a. s. l.; 30 Sep. 2009; Londt, Jason, Dikow, Torsten leg.; thick woody fynbos; NMSA -DIP-92350, NMSA • 1 ♂ Springbok, 8 km NW, road to Nababeep; 29°37'16"S, 017°49'59"E; 985 m a. s. l.; 30 Sep. 2009; Londt, Jason, Dikow, Torsten leg.; thick woody fynbos; NMSA -DIP-92351, NMSA South Africa – Western Cape • 1 ♂ Brenton on Sea; 34°04'20"S, 023°01'07"E; 10 Dec. 1979; Londt, Jason, Stuckenberg, Brian leg.; dune and hillside vegetation; NMSA -DIP-050742, NMSA • 1 ♂ Brenton