Onthophagus transisthmius Howden & Young 1981

Onthophagus transisthmius Howden & Young, 1981 Fig. 62B Onthophagus transisthmius Howden & Young, 1981: 106 (original description). Type locality: Panama: Colón: Colón: Gamboa. Name-bearing type: holotype (USNM), not examined. Onthophagus (Onthophagus) transisthmius — Zunino & Halffter 1997: 161 (list). — Escobar 2000: 209 (list for Colombia). — Medina et al. 2001: 140 (checklist for Colombia). — Ratcliffe 2002: 17 (checklist for Panama). — Pulido-Herrera & Zunino 2007: 117 (catalogue). — Delgado & Curoe 2014: 65 (key, cited for Panama). — Chamorro et al. 2018: 97 (cited for Ecuador). — Rossini et al. 2018a: 547, 552–553, 563‒564, figs 2a‒d, h‒i, 5 (key, diagnosis, redescription, remark); 2018b: 10 (osculatii complex). — Chamorro et al. 2019: 198 (catalogue). Distribution Panama, Colombia, Venezuela, Ecuador, Peru, and Bolivia. Subregions of Venezuela Plains, Serrania de Perija, and Andes mountains. Literature records Rossini et al. 2018a: 552, 563 (Venezuela: Táchira, Portuguesa and Zulia); 2018b: 11 (two species reaching eastern Amazonia). — Chamorro et al. 2019: 198 (Venezuela).