Spilogona padlei Huckett 1965

Spilogona padlei Huckett, 1965 Figs 5 H–J Spilogona padlei Huckett, 1965: 228. Type-locality: “Padlei, Northwest Territories, 7.25.50 [i.e. 25.vii.1950], R. E. Duckworth ” [CANADA, NWT] [seen]. Spilogona perambulata Huckett, 1965: 228. Type-locality: “Richardson Highway mi. 236, Alaska, 6.26.51 [i.e. 26.vi.1951], J.R. McGillis ” [USA, Alaska] [seen]. Syn. nov. Spilogona padlei Huckett, 1965; Sorokina & Thidrikh, 2021: 225 (Chukotka AO). Notes. Huckett described Spilogona padlei and Spilogona perambulata in the same monograph (Huckett, 1965). Huckett indicated in the description (1965: 228): “The species padlei and perambulata have much in common. The former however may be distinguished from the latter and related forms by the presence of a fine ventral bristle on mid tibia.” The type material of Spilogona padlei and Spilogona perambulata (in Canadian National Collection) has been examined. The descriptions of both species and the male terminalia are the same. Both species have sternite 1 with fine hairs, anepisternum without interspatial seta, scutellum with 2 long downwardly-directed preapical setulae on upper border of declivities, and male mid femur with 1–2 stout pv setae at base. However, Huckett noted in the description of Spilogona padlei “… mid femur with 2 or 3 fine pv,..”, and “… mid femur with a few weak pv …” for Spilogona perambulata. The mid femur of Spilogona padlei really does have a row of weak short pv setae in addition to stout pv seta. Huckett probably did not notice the stout setae because the mid legs are tucked close to the thorax. The mid tibia of Spilogona padlei is actually unlike that of Spilogona perambulata by possessing very short v setae. Since only 6 specimens (5 males and 1 female) of Spilogona padlei have been described from one locality (NWT, Padlei), it can be assumed that the short v setae are a variation of this character in one generation and not enough to describe a separate species. The male terminalia of Spilogona padlei and Spilogona perambulata from different generations is the same (Fig. 5H). The sternite 5 is especially distinctive because the two small tubercles at middle are visible in dry specimens (Fig. 5I). The females of both species are also identical; the ovipositor is especially distinctive, with large coarse hook-shaped spinules on the postgenital plate (Fig. 5J). I therefore conclude that Spilogona perambulata Huckett, 1965 and Spilogona padlei Huckett, 1965 are both new synonyms, and as they were both published in the same work and on the same page I select as first reviser Spilogona padlei as the valid name for this species. Additional material examined. RUSSIA, Taymyr Peninsula: 1♂, Verkhnyaya Taimyra River, mouth of Logata River, willow forest in floodplain, 30.vii.1967, leg. K. Gorodkov (ZISP). Wrangel Island: 1♂ 7♀, lower reaches of Somnitel′naya River, 70º56’N 179º37’W, yellow pan trap, 7–10.vii.2019, leg. P. Kulemeev & O. Khruleva. Magadan area: 2♀, 120 km NNE Seimchan, Lyadinoi stream, 63°44’N 153°25’E, yellow pan trap, 10.vii.2019, leg. N. Tridrikh; 2♀, ~ 100 km NNE Seimchan, Nature Reserve “Magadansky”, 63°39’N 153°17’E, valley of Suksukan River, thickets of young willow, white pan trap, 13.vii.2019, leg. N. Tridrikh; 2♀, the same data, but yellow pan trap, 21–24.vii.2019; 1♀, the same data, but pebble beach, pan trap, 24.vii.2019; 1♀, 10 km N Evensk, Tainyi stream, 61°55’N 159°16’E, 25.vii.2020 leg. N. Tridrikh; 1♀, 10 km W Evensk, Malaya Garmanda River, 61°52’N 159°23’E, sweeping in grass, 01.vii.2020, leg. N. Tridrikh (SZMN except where stated otherwise). Distribution. PALAEARCTIC: Russia (Arctic Siberia, NE of Far East). NEARCTIC: NWT, Alaska, Yukon, Quebec.