Paraputo Laing 1929

GenusParaputoLaing Type species. Lachnodiopsis szemaoensis Borchsenius, by original designation. Generic diagnosis (adapted from Williams (2004). Body of adult female broadly oval to rounded, membranous. Antennae each with 6 to 8 antennomeres. Legs well developed, generally robust, femur often about twice as wide as tibia. Combined length of tibia and tarsus usually shorter than that of trochanter and femur. Translucent pores commonly present on hind coxae, sometimes also on hind femur and tibia, and occasionally also on coxae of middle legs. Claws stout and lacking a denticle. Labium typically elongate, usually longer than clypeolabral shield. Anal ring generally located on dorsum at a distance equal to or greater than its length from apex of abdomen and bearing six or more setae. Circulus well developed or absent. Cerarii numbering 11 to 18 pairs; those on anal lobes and posterior abdominal segments each usually containing multiple conical setae, sometimes accompanied by normal setose setae that are often of similar basal width to conical setae, along with clusters of trilocular pores. In rare cases, cerarii each reduced to only two conical setae; often with intermediate cerarii or conical setae present also, thus forming a continuous row of conical setae around dorsal margin, frequently accompanied by a continuous band of densely packed trilocular pores; conical setae rarely present on ventral margin. Ostioles prominent, situated away from body margins, each wide, with inner lip edges sclerotized; each lip often with abundant trilocular pores, rarely with few, and bearing few to numerous setae. Eyes present. Spiracles large and conspicuous. Dorsum with setae usually numerous, either minute and stiff or longer and flagellate, often with longer setae flanking anal ring. Trilocular pores abundant on both surfaces. Multilocular disc pores and oral collar tubular ducts absent. Venter usually with flagellate setae, sometimes resembling those on dorsum. Cisanal and obanal setae usually apparent, sometimes each long and stout and displaced onto dorsum posterior to anal ring. Oral collar tubular ducts present, usually of only 1 size, either 1.5–2.0 times wider than a trilocular pore (referred to as large type), or as wide as, or narrower than a trilocular pore; but sometimes 2 or 3 different sizes present. Tubular ducts sometimes present across medial area and in marginal groups on abdomen, sometimes also present between antennal bases; present or absent between anal lobes. Anal lobes each either membranous or with various degrees of sclerotization, at times sclerotization occupying most of lobe, but never with a distinct anal lobe bar; occasionally ventral margins of some anterior abdominal segments also sclerotized (Williams and Granara de Willink 1992; Williams 2004; Danzig and Gavrilov 2010).