Capulus ungaricus

Capulus ungaricus (Linnaeus, 1758) (Fig. 10C 1, C 2) Patella ungarica Linnaeus, 1758: 782. Capulus (Capulus) hungaricus [sic] – Domènech & Martinell 1982: 50, fig. 15. Capulus ungaricus – Landau et al. 2011: 14, pl. 4, fig. 7. — Chirli & Linse 2011: 63, pl. 14, fig. 5. — Van Dingenen et al. 2016: 123, pl. 4, fig. 4. — Chirli & Forli 2017: 54, fig. 15d. For more, see synonymy list in Landau et al. (2011) and Van Dingenen et al. (2016). MATERIAL AND DIMENSIONS. — Maximum height 25.2 mm, width 11.9 mm (incomplete). — RGM.1364918 (4), leg. WG; RGM.1364997 (18), leg. ACJ; RGM.1365102 (57), leg. AWJ; RGM.1365200 (1), leg. AWJ; RGM.1364998 (1), leg. ACJ. SPECIES CHARACTERISATION. — Medium sized capuliform shell with ovate aperture, convex elevated in profile with coiled, recurved apex displaced slightly to the right overhanging posterior margin. Protoconch of just under two smooth whorls with small nucleus, rapidly expanding last whorl sculpture of fine axial ribs. U-shaped internal muscle scar. DISTRIBUTION. — Middle Miocene: Paratethys, Poland (Bałuk 1975), Vienna (Hörnes 1856); Proto-Mediterranean, Italy (Sacco 1896b); Atlantic, Aquitaine Basin, France (Cossmann & Peyrot 1919), Loire Basin, France (Glibert 1949). — Upper Miocene: Proto-Mediterranean, Italy (Sacco 1896b). — Lower Pliocene: NSB, Coralline Crag, England (Wood 1848; Harmer 1916), Kattendijk Formation, Belgium (Glibert 1958; Marquet 1998); Atlantic, NW France (Van Dingenen et al. 2016), SW Spain (Landau et al. 2011); central Mediterranean, Italy (Chirli 2008). — Upper Pliocene: NSB, Red Crag, England (Wood 1848; Harmer 1916), Oorderen Sands, Belgium (Glibert 1958; Marquet 1998); Atlantic, Portugal (Zbyszewski 1959; Silva 2001); western Mediterranean, S Spain (Landau et al. 2004), France (Chirli & Richard 2008); central Mediterranean, Italy (Sacco 1896b; Cavallo & Repetto 1992). — Upper Pliocene-Pleistocene: NW France (Brébion 1964). — Lower Pleistocene: Atlantic, Selsoif, NW France (this paper); central Mediterranean, Italy (Cerulli-Irelli 1914; Brambilla & Galli 1988; Di Geronimo & La Perna 1997; Chirli & Forli 2017); eastern Mediterranean, Rhodes Island (Chirli & Linse 2011). — Upper Pleistocene: western Mediterranean, Spain (Domènech & Martinell 1982). Today this species occurs in the Atlantic from Iceland to West Africa and North America to Florida, Mediterranean. It is not present in the North Sea (Fretter & Graham 1981). It inhabits in the sublittoral zone to 850 m depth, attached to stones and shells (Poppe & Goto 1991) REMARKS The Selsoif specimens have a protoconch of about two whorls, similar to that illustrated by Fretter & Graham (1981: fig. 225). For further discussion see Van Dingenen et al. (2016).