Siphonaria savignyi Krauss 1848

Siphonaria savignyi Krauss, 1848 (Figs 14J–L, O) Siphonaria savignyi Krauss 1848: 61.— Hanley 1858b: 152; Paetel 1873: 117; 1875: 92; 1883: 178; Tomlin 1927: 291; Bouchet & Danrigal 1982: 15; White & Dayrat 2012: 67. Siphonaria (Siphonaria) savignyi — Hubendick 1945: 72; 1946: 55, pl. 4, figs 25–29. Siphonaria (Siphonaria) laciniosa forma savignyi — Morrison 1972: 56–58, 60, 59, figs 3, 4. Siphonaria kurracheensis — Barash & Danin 1972: 329, fig. 14; 1977: 98 (not S. kurracheensis Reeve, 1856). Siphonaria savignyi Krauss, 1848 — Coan & Kabat 2017: 205. Material examined. Type material. Lectotype of S. savignyi Krauss, 1848, present designation, type locality unknown, likely Red Sea; collector and date unknown (Savigny collection) (IM 2000-35936, Fig. 14J). Paralectotypes, same data as lectotype (MNHN IM 2000- 35935 d, Fig. 14K; MNHN IM 2000-35934 d, Fig. 14L). Taxonomic remarks. In the original description, Krauss (1848) attributed this name to Philippi. However, Krauss is author of the name, and not Philippi in accordance with Art. 12.2.6 of the Code (see also Bouchet & Danrigal, 1982: 15; Kabat & Coan, 2017: 205). The new name S. savignyi has been made available by Krauss (1848) through providing a bibliographic reference to pl. 1, fig. 1 in the work of Savigny (1817). Hubendick (1946: 55) incorrectly attributed the name to an alleged publication by Philippi (1826) entitled “Gasteropodes et Qoquille in Savignyi: Descriptions de l’Egypte”. However, this title is that of Savigny’s (1817) work and no published work was authored by Philippi under this title (Kabat & Coan, 2017). Shell (Figs 14J–L; Table S9). Small to medium sized (max sl mean = 41 mm, SD = 2.2 mm, n = 3), ovate; height medium to tall; apex offset weakly posterior and to left, apical sides strongly convex, protoconch direction undetermined; shell whorl dextral; exterior whitish, uneven without prominent radial colour bands, growth striae prominent in bands, shell thickness thick; rib count (mean = 49, SD = 2.9, n = 3); ribs fairly even in width, straight to wavy, ridges rounded narrow, slightly broaden and increasingly raised to shell edge, 8–10 spread prominent whitish primary ribs whitish; siphonal ridge formed by 3–5 primary ribs, flared up at shell edge; few finer secondary ribs, rib interstices darker, edge weakly scalloped and unevenly corrugated; interior shell margin white, short dark brown rays on shell lip, align under rib interstices, margin thickened white, siphonal groove distinct; same colour as mottled reddish to dark brown spatula; ADM scar distinct, darker brown, CMS slightly convex; thickening and whitening of shell lip occurs (Fig. 14J). Reproductive system and Spermatophore. unknown. Comparative remarks. Figured shells from Port Fenfick and Gulf of Suez, Red Sea, and the description in Hubendick (1948: 55, pl. 4, 26–29) and figured shells from Israel in Morrison (1972: 60, figs 3–4) exhibit a morphology consistent with the types (Figs 14J–L). No material has been available to study the anatomy and/or mitochondrial phylogenetics of this species. Distribution and habitat. So far recorded from the Red Sea and the Mediterranean coast of Israel.