Columbellaria corallina

Columbellaria corallina (Quenstedt, 1852) Plate 10: figs 12–15 2017 Columbellaria cf. corallina (Quenstedt, 1852) – Werner, Nützel and Nose: 32, fig. 3 A – C. 2019 Columbellaria corallina (Quenstedt, 1852) – Gründel, Keupp and Lang: 133, pl. 9, figs 11–17 (here more chresonymy and synonymy). 2019 Columbellaria sp. 1 – Gründel, Keupp and Lang: 136, pl. 9, fig. 18, pl. 10, fig. 1. Material. 3 specimens, SNSB-BSPG 2016 XXI 1859 –1861. Description. Shell egg-shaped, higher than wide, with distinctly elevated, slender, gradate spire, consisting of more than 5 whorls; illustrated specimen (Plate 10: figs 14, 15) 21 mm high; already first preserved whorls angulated forming edge that later becomes a keel; keel separates oblique ramp and vertical abapical whorl face; ramp ornamented with several very weak spiral cords over several whorls; spiral cords no longer visible on last two spire whorls; instead, formation of subsutural spiral cord covered with small tubercles and a stronger spiral cord, also with nodes, between subsutural cord and keel; whorl face of early teleoconch with axial ribs (approximately 10 on last spire whorl); intersections of axial ribs and spiral cords nodular; all mentioned spiral cords continue onto body whorl without interruption; axial ribs disappear at transition from spire whorls to body whorl and keel is weakened into a spiral cord of equal strength as in other spiral cords; transition from convex whorl face to strongly convex base evenly rounded; body whorl covered with approximately 15 nodular spiral cords of approximately same strength; spiral cords on whorl face slightly more distant from each other than those on base; aperture with widened outer lip and spine-like protrusions where spiral cords are situated; aperture elongated with anterior and posterior siphonal canals (see Hägele 1997; Werner et al. 2017). Remarks. The spire of the present specimens combines characteristics of Columbellaria corallina sensu Gründel et al. (2019) (the last whorl is approximately 10 mm wide; distinct spiral cord between adapical suture and keel) with that of Columbellaria sp. 1 sensu Gründel et al. (2019) (strong keel on whorls until end of spire, stronger nodes on keel). It is possible that all forms described Gründel et al. (2019) and described here are variants of a single species. Due to the lack of more extensive and better-preserved material, these forms are summarized herein under the name Columbellaria corallina (Quenstedt, 1852). Columbellaria rara Gründel et al. (2022) has significantly fewer spiral cords on the body whorl and these are more bulging. Zittelia picteti Gemmellaro, 1870 has a wider and more convex body whorl, the spiral cords are at least as wide as the spiral furrows between them and the spire whorls are not keeled. Columbellaria victoria Guirand & Ogérien, 1865 and sensu Loriol in Loriol and Bourgeat (1886–1888) has spiral cords of equal strength on the body whorl, which are at least as wide as the spiral furrows between them. Its body whorl is more convex and it lacks a subsutural row of small nodes. Columbellaria denticulata Zittel, 1873 has more nodes on the last spire whorl and lacks subsutural row of small nodes is missing there; the outer lip of the body whorl lacks furrows.