Endemulia Brown 2024, new genus

Endemulia Brown, new genus LSIDurn:lsid:zoobank.org:act: 72968E15-C869-40EF-83FE-EDF63BE797E4 Type species: Endemulia urrai Brown, new species. Remarks and Diagnosis. Endemulia is described for a single new species that is superficially somewhat similar to Accuminulia but with male genitalia similar to those of Chapoania. In forewing length and pattern, Endemulia urrai is reminiscent of Accuminula buscki Brown, with rather ill-defined pattern elements of gray and pale brown on a paler ground color, and a slender, short, dark, oblique dash near the end of the discal cell. However, the genitalia show little similarities to those of A. buscki. Instead, the long, free, spiny, terminal end of the sacculus is similar to that of Chapoania, to which I initially assumed the species should be assigned. However, adults of Endemulia are dissimilar to those of Chapoania in forewing length and pattern (Figs. 13, 14), and in the female genitalia (Figs. 45, 46). Hence, a new genus is proposed for this species. Description. Head: Vertex rough-scaled, scales directed anteriorly; upper frons rough-scaled, scales of lower frons shorter, more appressed to surface; antenna ca. 0.5 length of forewing, simple, with one row of scales per flagellomere, sensory setae ca. 0.5 times width of flagellomere in male, antenna more slender in female, with shorter and sparser sensory setae; ocellus small; chaetosemata present; labial palpus with scaling expanded distally on second segment, third segment exposed, combined length of all segments ca. 2.5 times diameter of compound eye in both sexes. Haustellum well developed, presumably functional. Thorax: Dorsum smooth-scaled, metathoracic tuft present. Legs unmodified, without hairpencil or other male secondary scales. Forewing length 7.0‒8.0 mm (n = 4) in males, 8.0‒ 9.5 mm (n = 2) in females, length ca. 2.4 times width, slightly expanded apically; costa slightly and evenly arched throughout, male costal fold absent; termen straight, oblique; all veins present and separate, chorda absent, M-stem present, R 4 to costa, R 5 to termen. Hindwing with all veins present and separate; CuP weak at margin; male with short, slender patch of secondary scales along lower margin of discal cell in basal 0.5 of underside of wing; frenulum with one bristle in male, three in female. Abdomen: Male genitalia (Fig. 34) with tegumen slightly narrowed dorso-posteriorly, vinculum U-shaped; uncus simple, rod-like, curved subbasally, with a pair of long, slender, straight setae from dorsum subbasally; socius sclerotized basally into a bilobed process with membranous, hairy portion between lobes; gnathos arms joined at 0.5 length into slender, apically pointed terminal plate, with fine spines from venter; transtilla with subtriangular, tonguelike, strongly spined, median process; valva comparatively short, costa nearly straight, sacculus strongly sclerotized, confluent with ventral margin of valva in basal 0.5, terminating in a long, free, sightly curved, densely spined process extending across face of valva; juxta a subtriangular plate; phallus relatively short and stout, slightly curved ventrad in distal 0.2 with pointed ventral tip; vesica with linear patch of 10‒12 short, non-deciduous cornuti. Female genitalia (Fig. 46) with papillae anales rather slender, ca. 4.5 times as long as wide, apophyses slender, posteriores ca. 2 times as long as anteriores; segment eight with a pair of sclerotized triangular plates with apices approaching midline of venter; sterigma complex, with two paired sublateral arms, each pair joined mesally by a semicircular plate in parallel fashion; ductus bursae broad, strongly sclerotized, slightly shorter than corpus bursae, gradually broadening into corpus bursae; corpus bursae with numerous parallel wrinkles throughout; signum absent. Etymology. The genus name is a combination of the words “endemic” and the genus name “ Eulia.”