Euura tillbergi

264 Euura tillbergi (Malaise, 1921) Figs 284–289, 489–491 Amauronematus tillbergi Malaise, 1921b: 125–126. Lectotype designated below. Amauronematus tillbergi var. foveifrons Malaise, 1921b: 126–127. Lectotype designated below. Synonymy by Lindqvist (1941b). Amauronematus latisulcatus Malaise, 1931b: 45–46. Syn. nov. Amauronematus pacificus Malaise, 1931b: 47–48. Lectotype designated below. Secondary homonym of Pontania pacifica Marlatt, 1896 [Euura pacifica (Marlatt, 1896)]. Syn. nov. Amauronematus mcluckieae Benson, 1935: 31–32. Mandatory correction of original orthography mcluckiei. Syn. nov. Amauronematus rex Benson, 1948b: 32. Syn. nov. Amauronematus obscurus Lindqvist, 1962b: 11–12. Secondary homonym of Nematus obscurus Norton, 1861 [Euura obscura (Norton, 1861)]. Syn. nov. Amauronematus pustulatus Lindqvist, 1962b: 12–13. Syn. nov. Euura tranquilla Vårdal & Prous, 2014 in Prous et al. 2014: 52. Replacement name for Amauronematus pacificus Malaise, 1931. Diagnosis Females (4.5–6.5 mm) with mostly or completely black abdomen (and thorax) can be distinguished from the other species in the vittata group. Paler females are most similar to those of E. hulteni, E. hedstroemi, E. minivittata sp. nov., and might not always be distinguishable. Euura hulteni typically have dorsally and ventrally black-lined metafemur (similar in E. minivittata), while the metafemur is basally black in E. tillbergi. Euura minivittata has a darker pterostigma and metafemur than in pale specimens of E. tillbergi. The pterostigma of Euura hedstroemi is typically posteriorly dark-lined and the metatibia with a dark spot at basal third of tibia, but pterostigma and metatibia can be (almost) uniformly pale, making separation from paler specimens of E. tillbergi unreliable. Male penis valves are most similar to E. krausi and paler specimens might not be distinguishable from this species. In E. tillbergi valvula 3 is rarely triangular, without a posterior invagination (ZMUO.031718). Male 4.0–6.0 mm. Type material examined Lectotype Amauronematus tillbergi, here designated SWEDEN – Torne Lappmark • ♀; Torneträsk; 68.22° N, 19.74° E; 400 m a.s.l.; R. Malaise leg.; NHRS, NHRS-HEVA000003942. Lectotype Amauronematus tillbergi var. foveifrons, here designated SWEDEN – Torne Lappmark • ♀; Torneträsk; 68.22° N, 19.74° E; 400 m a.s.l.; R. Malaise leg.; NHRS, NHRS-HEVA000003943. Notes Best matches: DEI-GISHym84434 (colouration), DEI-GISHym84433 (size), DEI-GISHym83702 (lancet). Holotype Amauronematus latisulcatus RUSSIA – Kamchatka oblast • ♀; Klutchi [Klyuchi]; 56.32° N, 160.85° E; R. Malaise leg.; NHRS, NHRS-HEVA000006415. Lectotype Amauronematus pacificus, here designated RUSSIA – Kamchatka oblast • ♀; Petropavlovsk; 55.06° N, 158.65° E; R. Malaise leg.; NHRS, NHRS-HEVA000006416. Holotype Amauronematus mcluckieae UNITED KINGDOM – Scotland • Cairngorm Mts, Cairngorm; 57.09° N, 3.69° W; above 910 m a.s.l.; 27 Jun. 1934; R.B. and J.E. Benson leg.; BMNH, B.M.TYPEHYM.1.642. Holotype Amauronematus rex UNITED KINGDOM – England • ♀; Buckinghamshire, Whaddon Chase; 51.83° N, 0.82° W; Mar. 1946; R.B. Benson leg.; reared ex larva on Salix; BMNH, B.M.TYPEHYM.1.685. Notes Best morphological match: DEI-GISHym83872. Was previously synonymized with hedstroemi Malaise, but colour characters of the rex holotype fit better with specimens of E. tillbergi from the Russian Far East (DEI-GISHym83872, sequenced). Mesoscutellum is not completely pale in any E. hedstroemi specimens studied. In addition, the type specimens of rex Benson were reared from Salix aurita and possibly from S. atrocinerea (mixed rearing), but E. hedstroemi seems to be restricted to S. phylicifolia. Holotype Amauronematus obscurus FINLAND – Uusimaa • ♀; Helsinge [Vantaa]; 60.30° N, 24.90° E; 25 May 1961; E. Lindqvist leg.; MZH, http://id.luomus.fi/GL.2750. Holotype Amauronematus pustulatus FINLAND – Kuusamo • ♀; Kitka; 66.24° N, 28.98° E; 5 Jul. 1958; M. Viitasaari leg.; MZH, http://id.luomus.fi/GL.2536. Host plants Salix phylicifolia, S. myrsinites, S. herbacea, and possibly S. aurita and S. atrocinerea Brot. if Amauronematus rex Benson really is a synonym of E. tillbergi. Genetics COI Based on 35 specimens, maximum within-species distance is 3.28% and the nearest neighbour, diverging by a minimum of 0%, is Euura krausi. Nuclear Based on 31 specimens, maximum within-species distance is 1.7% (1.08% based on haplotypes of individual females). The nearest neighbour, diverging by a minimum of 0.12%, is Euura krausi. Distribution and material examined Palaearctic. Specimens studied are from Finland, Norway, Russia, and Sweden.