Euura papillosa ZMUO.

198 Euura papillosa group Bifid claws, short and narrow valvula 3 in dorsal view, pale elongate pterostigma, extensively pale body and structure of lancet (radix about as long as lamnium, setae long, basalmost annulus roughly 0.5 times as long as broad, basal sutures in dorsal part inclined basally and apical sutures apically) distinguish the females from those of the other Euura. Males are also extensively pale. Penis valves can be distinguished from the other Euura. Penis valve length 1.1–1.3 mm, paravalva broader apically than basally, ventral and anterior margins of paravalva distinctly angled, straight and rather narrow and long valvispina, symmetrically narrowing and round pseudoceps at apex, ventroapical margin of paravalva broad, inclined basally and without distinct invagination between paravalva and valvispina, basal third of valvar strut merging with ventral margin of paravalva. Euura papillosa group females lay eggs on the underside (abaxial) surface of leaves (Urban 2001), in contrast to most other Euura that lay eggs inside the leaf or petiole tissue. This habit of egg laying is shared with E. spiraeae (Robbins 1927; Hara et al. 2022) and the ribesii group (Viitasaari 1980) with a similar type of lancet. Euura hypoxantha and E. jugicola (myosotidis group) also have a similar type of lancet, but egg laying habits of these species have not been described to our knowledge. Two species are recognised.