Pristimantis yonke Chávez, Aznaran, Wong, Victoriano-Cigüeñas, García-Ayachi, Valencia-Málaga, Ormeño, Gulman, Sumiano-Mejía, Thompson & Catenazzi, 2025, sp. nov.

Pristimantis yonke sp. nov. Type material. Holotype • Adult male CORBIDI 26399 (Figs 11 A ‒ D, 12 A ‒ E) from Laguna Negra trail, Ayabaca Province, Piura Region, Peru (4°55'52.58"S, 79°29'8.41"W; 2933 m a. s. l.) collected on 6 September 2023 by Juan D. Valencia-Málaga and Germán Chávez. Paratypes • Adult female CORBIDI 26398, male CORBIDI 26402 collected with the holotype, • adult males CORBIDI 26413 ‒14 collected at the type locality on 9 and 10 September 2023, respectively by Juan D. Valencia-Málaga, and Germán Chávez. • Adult female CORBIDI 26929 from Bosque de Ramos, Ayabaca Province, Piura Region, Peru (4°42'25.75"S, 79°27'51.24"W; 2949 m a. s. l.), collected on 9 July 2024 by Wilmar Aznaran, Karen Y. Victoriano-Cigüeñas, Jesús R. Ormeño, Ronal Sumiano-Mejía, and Germán Chávez (Fig. 13 A ‒ H). Referred specimens. Juvenile CORBIDI 26403 from the type locality; juveniles CORBIDI 26924 ‒25 from Bosque de Ramos, Ayabaca Province, Piura Region, Peru (4°42'25.75"S, 79°27'51.24"W; 2949 m a. s. l.), collected on 9 July 2024 by Wilmar Aznaran, Karen Y. Victoriano-Cigüeñas, Jesús Ormeño-Benavides, Ronal Sumiano-Mejía, and Germán Chávez. Diagnosis. Pristimantis yonke sp. nov. is a small species with broad flat head, which may be distinguished by the following combination of traits: (1) skin on dorsum finely tuberculate without crests or ridges on scapular region, incomplete and low dorsolateral fold occasionally present; skin on venter areolate; discoidal fold present; thoracic fold absent; skin on flanks as on dorsum; longitudinal middorsal fold evident; (2) tympanic membrane and tympanic annulus evident, its length about 50 % of the length of eye; supratympanic fold present, going from the distal edge of tympanum to the level of the insertion of forelimbs; (3) snout short, dorsoventrally compressed, acuminate in dorsal view, protruding and inclined posteroventrally in profile, rostral papillae present; canthus rostralis concave; (4) upper eyelid having small, low rounded tubercles, about 70 % IOD in males, 50 % in females; cranial crests absent; (5) dentigerous processes of vomers absent; (6) males with a subgular vocal sac and vocal slits; (7) Finger I slightly shorter than Finger II; discs on fingers broadly expanded, elliptical (Fig. 12 C); (8) fingers having lateral fringes; (9) ulnar tubercles absent; (10) heel and tarsus lacking conical tubercles, only low rounded tubercles present; (11) inner metatarsal tubercle elliptical, elevated, about 3 times the size of round outer metatarsal tubercle; supernumerary tubercles low; (12) toes with lateral fringes; basal webbing on feet; Toe V longer than Toe III (disc on Toe III slightly surpasses distal edge of the penultimate subarticular tubercle on Toe IV, disc on Toe V clearly surpasses the distal edge of the distal subarticular tubercle on Toe IV) (Fig. 12 D); (13) in life, dorsum dark brown to cinnamon brown, with or without dark blotches; flanks same color as dorsum; head having dark supratympanic fold, pale interorbital bar occasionally present; groins, axillae and posterior surface of thighs creamy white to dark brown with or without yellow mottling; throat cream, creamy yellow or yellow with or without dark brown reticulations; belly creamy white or creamy yellow with dark dots, ventral surfaces of thighs yellowish pink or yellow with dark dots; iris gold with thin black reticulations, and a broad horizontal reddish brown streak (Fig. 11 A – D); (14) average SVL in adult males: 19.6 ± 0.3 mm (19.3–20.1 mm; n = 4); in females: 30.6 ± 2.4 mm (28.9–32.3 mm; n = 2). Comparisons. Pristimantis yonke sp. nov. is closely related, and also similar in appearance, to P. morlaco Sánchez-Nivicela, Toral-Contreras and Urgiles 2022, however, the new species can be differentiated by having an acuminate snout from dorsal view (vs subacuminate), inclined posteroventrally from profile (vs acuminate), palmar tubercle oval (vs “ U ” shaped), lacking dentigerous processes of vomers (vs present, oblique) and males being smaller with a SVL of 19.3–20.1 mm (vs 24.1–26.6 mm). The new species shares with members of the P. lacrimosus group (Duellman and Lehr 2009; Rivera-Correa and Daza 2020; Carrión-Olmedo and Ron 2021; Castillo-Urbina et al. 2023) and members of the P. colodactylus subgroup (Székely et al. 2016) the dorsoventrally compressed body and the flat, broad, pointed head. From members of the P. lacrimosus group, Pristimantis yonke sp. nov. can be easily distinguished by having a dorsum predominantly brown (vs olive green, green, yellow, gold or orange in P. achupalla, P. acuminatus, P. bromeliaceus, P. ecuadoriensis, P. enigmaticus, P. eremitus, P. galdi, P. jorgevelosai, P. lacrimosus, P. latericius, P. limoncochensis, P. loeslein, P. mendax, P. moro, P. nankints, P. olivaceus, P. omeviridis, P. ornatissimus, P. padiali, P. pardalinus, P. petersi, P. petersioides, P. pseudoacuminatus, P. rhodoplichus, P. romeroae, P. royi, P. schultei, P. tantanti, P. tayrona, P. urani, P. waoranii, and P. zorro). Only P. amaguanae, P. aureolineatus, P. mindo, P. nyctophylax, P. pluvialis, P. pulchridormientes, P. subsigillatus and P. zimmermanae have a brown dorsum among their color patterns, but the new species is distinguishable by having a snout protruding in profile (vs acuminate in P. nyctophylax and P. zimmermanae, rounded in P. mindo and P. pluvialis), having a rostral papillae on the tip of the snout (vs absent in P. aureolineatus, P. mindo, P. nyctophylax, P. pulchridormientes, and P. subsigillatus), having skin on dorsum finely tuberculated (vs. shagreen in P. amaguanae, P. aureolineatus, and P. pulchridormientes; smooth in P. mindo, P. pluvialis, P. subsigillatus, and P. zimmermanae), lacking vomerine teeth (vs present in P. aureolineatus, P. mindo, P. nyctophylax, and P. subsigillatus), and lacking conical tubercles (vs present on heels in P. amaguanae, P. nyctophylax, and P. subsigillatus). From members of the P. colodactylus subgroup, P. yonke may be differentiated by the combination of the following features (characters for other species in parenthesis): having an average SVL of 20.7 mm in males and 30.6 mm in adult females (vs 16.9 mm in males and 20.3 in females of P. colodactylus, 20.4 mm in males and 22.5 in females of P. matildae, 16.8 mm in males and 22.1 in females of P. muranunka), snout acuminate in dorsal view (vs rounded in P. muranunka), skin on dorsum tuberculate (vs areolate in P. colodactylus), longitudinal middorsal fold present (vs absent in P. colodactylus and P. muranunka), tympanic membrane and annulus present (vs absent in P. colodactylus and P. matildae), conical tubercles on upper eyelid and heels absent (vs present in P. matildae), larger feet size being about 44 % of the SVL (vs 36 % of SVL in P. colodactylus, 38 % of SVL in P. muranunka, 40 % in P. matildae), and iris coloration gold (vs bronze in P. colodactylus, reddish bronze in P. matildae and P. muranunka). Moreover, Pristimantis yonke is genetically related to Pristimantis cf. cajamarcensis (KU 217845). But it can be distinguished by being smaller, with males reaching up to 20.1 mm of SVL (vs males up 24.1 mm of SVL in P. cf. cajamarcensis), having dorsum tuberculate (vs shagreen in P. cajamarcensis), lacking dentigerous processes of vomers (vs present in P. cf. cajamarcensis), and lacking red or orange blotches on groins and posterior surface of thighs (vs present in Pristimantis cf. cajamarcensis, sensu Lynch 1979). Description of the holotype. An adult male (CORBIDI 26399). Measurements (in mm): SVL 19, 6.3; TL 10.2; FL 8.6; HL 7.3; HW 7.7; ED 2.6; TD 1.2; IOD 2.6; EW 1.9; IND 1.4; EN 2.4; FeL 9.5. Head wider than long, narrower than body; snout acuminate in dorsal view, protruding in profile, having a rostral papilla on the tip of the snout (Fig. 12 E); cranial crests absent; nostrils slightly protuberant, directed anterolaterally; canthus rostralis slightly concave in dorsal view, rounded in cross section; loreal region slightly concave; upper eyelid lacking conical tubercles, only small rounded present; tympanic annulus present, visible through the skin, its upper and posterolateral edge concealed by supratympanic fold; tympanic membrane visible; postrictal tubercles absent. Choanae median, ovoid, non-concealed by palatal shelf of maxilla; dentigerous processes of vomers absent; tongue slightly longer than wide, not notched, posterior half free; vocal slits slightly curved, positioned at posterior half of mouth floor in between tongue and margin of jaw; medium sized vocal sac. Dorsal surfaces of body and flanks finely tuberculate; skin on chest and belly areolate, that on throat shagreen, ventral surfaces of limbs shagreen, ventral surfaces of thighs coarsely areolate; discoidal fold present, thoracic fold absent. Ulnar tubercles absent; nuptial pads absent; outer palmar tubercle ovoid low, as large as ovoid thenar tubercle; subarticular tubercles low, rounded; large supernumerary tubercles at base of fingers, distinct; fingers bearing lateral fringes; Finger I shorter than Finger II; discs on fingers expanded and rounded; pads on fingers surrounded by circumferential grooves on all fingers (Fig. 12 C). Hindlimbs slender; dorsal surfaces of hindlimbs finely tuberculate; posterior surfaces of thighs smooth, ventral surfaces of thighs coarsely areolate; heel bearing low rounded tubercle; outer and inner edge of tarsus lacking tubercles; inner metatarsal tubercle elliptical, elevated, 1.5 × the size of oval outer metatarsal tubercle; plantar surface with small, low and rounded supernumerary tubercles; subarticular tubercles prominent, rounded; toes bearing lateral fringes; basal webbing between toes III and IV, and IV and V; discs on toes slightly smaller than those on fingers, expanded and rounded; all toes having pads surrounded by circumferential grooves; relative lengths of toes: I <II <III <V <IV; Toe V longer than Toe III (disc on Toe III reaches the distal edge of penultimate subart