Xymmer muticus

Xymmer muticus (Santschi, 1914) Figure 1A–C; 2A–C; 3A–C; 4A–E. Stigmatomma (Xymmer) muticum Santschi, 1914 d: 311, fig. 2 (w.) NIGERIA. (specimen images examined) Combination in Xymmer: Wheeler, 1922: 758. In Amblyopone: Brown, 1960 a: 168. In Xymmer: Yoshimura & Fisher, 2012 a: 19. Redescription—worker Figures 1A–C. Material examined (n = 14): Ivory Coast: Montagnes: Site 07 (Taï N. P.) 200 m, 5.84598, -7.34815, Primary Forest. Gómez, K., Kouakou, L. 13/11/2019, Hand, ex. rotten log; 1 worker on pin KGCOL03192, RBINSFOR1000801 [RBINS] • same data; 1 worker on pin KGCOL03959, RBINSFOR1000804 [RBINS] • same data; 1 worker on pin KGCOL03960, RBINSFOR1000805 [RBINS] • Site 07 (Taï N. P.) 200 m, 5.84598, -7.34815, Primary Forest. Gómez, K., Kouakou, L. 13/11/2019, Hand, ex. rotten log; 1 worker in ethanol KG04106, 2 workers on pin KGCOL03165 [KGAC] • Ivory Coast: Montagnes: Site 04 (Taï N. P.) 200 m, 5.83093, -7.344, Primary Forest. Gómez, K., Kouakou, L. 10/11/2019, Hand, Ex soil; 1 worker in ethanol KG04078, 1 worker on pin KGCOL03166 [KGAC] • same data; 2 workers on pin KGCOL03165 [KGAC] • Site 07 (Taï N. P.) 200 m, 5.84598, -7.34815, Primary Forest. Gómez, K., Kouakou, L. 13/11/2019, Hand, ex. rotten log; 2 workers in ethanol KG04341 [KGAC] • Ivory Coast: Oumé-Goulikao, Foret. Kone, M. 31/03/2009, M1 (0-15 cm); 1 worker on pin, RBINSFOR002633 [RBINS] • same data, Winkler, Leaf litter, 1 worker on pin, KGCOL03166 [KGAC]. Measurements (n = 3): HW 0.51–0.52; HW₂ 0.49–0.51; HL 0.57–0.59; MBL 0.36–0.36; SL 0.35–0.40; WL 0.82–0.83; ML 0.79–0.81; PNW 0.27–0.34; HFL 0.38–0.39; PL 0.41–0.42; PH 0.27–0.28; DPW 0.22–0.27; DPL 0.27–0.30; AIIIW 0.35–0.35; AIVW 0.35–0.38; AIIIL 0.24–0.28; AIVL 0.28–0.29; AVL 0.11–0.15; AVIL 0.07–0.10; AVIIL 0.17–0.26; TL 3.08– 3.20. Indices: CI 86.50–90.01; MI 69.13–71.54; SI 68.45–79.25; WI 241.54–302.63; HFI 72.72–77.67; AIIII 130.74– 161.86. Head In full-face view, head slightly longer than broad, subquadrate (CI 86.50–90.01); lateral head margins convex, reaching maximum width in anterior half; occipital margin weakly concave. Mandible linear, long (MI 69.13–71.54), widest medially; masticatory margin with 9 teeth, 5 of them being paired; first two basal teeth minute, unpaired; proceeding 5 teeth paired, triangular, subtly incrassate apically; subapical tooth unpaired, triangular; apical tooth large, sharply triangular. Eye absent. Anteromedial clypeal lobe, projecting into intermandibular space, straight to very subtly concave, lacking denticles. Frontal lobes concealing majority of antennal foramen. Frontal carinae minute, scrobes absent. Genal teeth absent. Scape failing to reach posterior head margin (SI 68.45–79.25), reaching its maximum width slightly apically to the mid-length. Antennal pedicel longer than wide, reaching maximum width apically; flagella segments I-IX wider than long and incrassate apically; flagella segment X longer than wides sharppointed apically. Mesosoma In profile view, dorsal outline gradually convex, highest at promesonotal articulation; posterior pronotal angle obtuse, propodeal dorsum gradually sloping towards declivity. In dorsal view, pronotum just slightly wider than propodeum; mesosoma narrowest at mesonotum and anterior region of propodeum; pronotal anterior margin convex, posterior margin concave; promesonotal articulation present, metanotal groove absent; propodeal declivity flat. In lateral view, lower propodeal lobes present. Lower-upper mesopleural suture absent; meso-metapleural suture present only between the lower portion of these sclerites. Anterolateral corner of the mesopleuron with a short blunt, anteriorly directly cuticular extension. Metapleural gland reservoir circular, occupying less than half of propodeal surface, located on the lower and posteriormost corner of propodeum in lateral view; propodeal spiracle circular, located below the median line of propodeum, separated from the reservoir by a distance about 1× its diameter. Propodeal lobe inconspicuous, semi-circular. Meso-metapleural suture present only between the lower portion of these sclerites. Profemora medially swollen. Tibiae roughly triangular, distinctly wider apically than at base. Metatibiae with pectinate comb and barbulate spur; mesotibiae with one spur. Pretarsal claws simple, lacking teeth on their inner margins. Metasoma In lateral view, petiole with distinct peduncle; anterior face straight; angle between petiole anterior face and dorsum obtuse; dorsum subtly convex across length; lacking posterior face. Posteroventral angle obtuse; anteroventral margin subtly concave, as long as posteroventral margin. Petiole spiracle circular; located just posterior to peduncle. In dorsal view, petiole narrow, distinctly longer than wide, lateral margins subtly convex.Anterolateral angle gradually curved. In ventral view, petiolar poststernite ovoid; narrow inverted U-shaped sulcus separating poststernite and laterotergite; ventrolateral junction marginated. In profile, ventral outline of gastral sternite of abdominal segment III distinctly more convex than corresponding tergite. Girdling constriction of abdominal segment IV conspicuous. Sting well-developed. Sculpture Head densely foveolate overall, with medial smooth central, longitudinal strip extending from clypeal lobes to vertexal margin. Mandible, scape and antennal segments punctulate, rendering a dull surface. Pronotum overall punctulated, mat, except for collar, with faint transverse strigulae, mesopleuron, metapleuron and propodeum including declivity finely striate. Visible portion of presternite III imbricate. Girdling constriction abdominal segment IV with areolate sculpturing across its width. Abdominal segments II-VII finely punctured with smooth and shining interspaces. Pilosity Whole body covered with appressed dense pubescence. Mandible masticatory margin lined with a series of erect, fine setae directed into inner mandibular space; mandible dorsum with subdecumbent setae of varying length. Anterior clypeal margin with a pair of long and numerous fine shorter setae that extend well into inter-mandibular space. Head dorsum, with pubescence directed medially. Frontal lobes with numerous, appressed setae radiating outwards. Meso-, metapleuron and declivity of propodeum mostly glabrous. Petiole peduncle glabrous. Gastral sternites with numerous, short subdecumbent setae as well as longer, erect setae; pygidium and hypopygium with a denser concentration of such setae, including the longest in metasoma, lacking spiniform setae. Legs covered with appressed pubescence. Colour Head, mesosoma, and metasoma orange yellow; mandible, scape, flagellum, and legs yellow. Description—gyne Figures 2A–C; 4B Material examined (n = 4): Ivory Coast: Montagnes: Site 07 (Taï N. P.) 200 m, 5.84598, -7.34815, Primary Forest. Gómez, K., Kouakou, L. 13/11/2019, Hand, ex. rotten log; 1 alate gyne on pin, KGCOL03193, RBINSFOR1000802 [RBINS] • same data; 1 dealate gyne in ethanol, KG04106 [KGAC] • Site 02 (Taï N. P.) 200 m, 5.83117, -7.3429, Primary Forest. Gómez, K., Kouakou, L. 08/11/2019, Winkler, Leaf litter; 1 dealate gyne on pin, KGCOL00224 [KGAC] • Site 06 (Taï N. P.) 200 m, 5.83458, -7.34638, Primary Forest. Gómez, K., Kouakou, L. 12/11/2019, Winkler, Leaf litter; 1 dealate gyne on pin KGCOL00282 [KGAC]. Measurements (n = 1): HW 0.55; HW₂ 0.54; HL 0.60; MBL 0.38; SL 0.37; WL 0.93; ML 0.89; PNW 0.38; HFL 0.42; PL 0.45; PH 0.30; DPW 0.25; DPL 0.30; AIIIW 0.41; AIVW 0.43; AIIIL 0.34; AIVL 0.37; AVL 0.20; AVIL 0.18; AVIIL 0.26; TL 3.71. Indices: CI 92.17; MI 69.44; SI 66.00; WI 237.77; HFI 75.59; AIIII 164.92. Head As in the worker, except: in full face view, eye present in posterior third of head; eye positioned laterally; with ca. 30 ommatidia and 7 across maximum width; eye barely surpassing lateral head margin. Ocelli present in posterior quarter of head, their width shorter than the distance between them. Masticatory margin of mandible with a maximum 8 teeth, 5 paired. Mesosoma With the usual gyne morphological features; mesosoma segmented, in dorsal view, pronotum visible; mesoscutum wider than long; notaular lines absent; parapsidal lines present, faintly impressed; scuto-scutellar suture present, narrow; metanotum present as a transverse strip, longer than wide; mesepimeral lobe indistinct; meso-metapleural sulcus narrow; metapleuron divided into upper metapleuron only, lower metapleuron fused to lateral propodeal surface. Metapleural gland reservoir large, almost reaching propodeal spiracle. Metasoma As in the worker, but metasomal segments III-VII enlarged. Wings Forewing; as in Figure 10. and Xymmer forewing description presented in Yoshimura and Fisher (2012). Hindwing; as in Figure 10. and Xymmer hindwing description presented in Yoshimura and Fisher (2012). Sculpture As in worker, except: mesoscutum and metanotum smooth and shining other than hair bearing punctures; meso-, meta- pleuron, and lateral face of propodeum with stronger striae covering the majority of each pleuron. Pilosity As in worker except for denser pubescence. Colour Head, mesosoma, metasoma, mandible, scape, flagellum, and legs light orange yellow. Description–male Figures 3A–C; 4C–E Material examined (n = 4): Ivory Coast: Montagnes: Site 07 (Taï N. P.) 200 m, 5.84598, -7.34815, Primary Forest. Gómez, K., Kouakou, L. 13/11/2019, Hand, ex. rotten log; 1 male on pin, KGCOL03192, RBINSFOR1000803 [RBINS] • Site 07 (Taï N. P.) 200 m, 5.84598, -7.34815, Primary Forest. Gómez, K., Kouakou, L. 13/11/2019, Hand, ex. rotten log; 3 males in ethanol, KG04106 [KGAC]. Measurements (n = 1): HW 0.45; HW₂ 0.31; HL 0.49; MBL 0.20; SL 0.12; WL 0.92; ML 0.94; PNW 0.52; HFL 0.43; PL 0.36; PH 0.26; DPW 0.21; DPL 0.18; AIIIW 0.29; AIVW 0.324; AIIIL 0.20; AIVL 0.20; AVL 0.20; AVIL 0.17; AVIIL 0.15; AVIIIL 0.11; TL 2.99. Indices: CI 91.08; MI 43.43; SI 26.50; WI 180.27; HFI 95.55; AIIII 135.38. Head In full-face view, head longer than broad (excluding eyes); posterior margin st