Sphaeroderma quadrimaculatum Chujo 1935

Sphaeroderma quadrimaculatum Chûjô, 1935 Fig. 14 E – L Host plant. Ranunculaceae: Clematis taiwaniana Hayata var. ryukiuensis Tamura. Leaf mine. Full-depth linear mine on mature leaf (Fig. 14 F – H, L). Frass is granular and minute, deposited linearly in two rows along both sides of the mine. The fully grown larva exits the mined leaf in early winter, falls to the ground, and pupates underground. Material examined. • 2 adults, Hedo, Kunigami, Okinawa Pref., 10-XI-2021 on Clematis taiwaniana var. ryukiuensis (Fig. 14 E); • 3 adults, Hedo, Kunigami, Okinawa Pref., 14-X-2000 (as larva on C. taiwaniana var. ryukiuensis), emerged on 21-XI – 10-XII-2000 (Fig. 14 F – H); • 2 adults, Tonaki Is., Shimajiri-gun, Okinawa Pref., 18-III-2020 (as larva on C. taiwaniana var. ryukiuensis), emerged on 10-V-2020 (Fig. 14 I – L).