Pupilla tetrodus

54. Pupilla tetrodus (O. Böttger, 1870) Distribution and year: Gokwe River 1869 (BÖttger 1870). Geographic range: Botswana, Lesotho, Namibia, South Africa, Zambia. Habitat: Terrestrial. Notes: The species is ubiquitous across Southern Africa, occurring from semi-arid regions of the Kalahari Desert to the coastal regions of the Eastern Cape (Herbert & Kilburn 2004). Type locality: Gokwe River (22°S 28°E, tributary of Motloutse River), Botswana (BÖttger 1870). Sources: BÖttger (1870: 46, as Pupa tetrodus); Connolly (1912: 185, as Jaminia tetrodus; 1939: 393); van Bruggen (1966 a: 109); Herbert & Kilburn (2004: 105). Conservation status: Not Evaluated.