Messapus Simon 1898

Updated key to the species of Messapus 1. Males (males of M. meridionalis Haddad & Mbo, 2015 and M. seiugatus Haddad & Mbo, 2015 unknown).............. 2 - Females............................................................................................. 7 2. Palp with single elongate slender RTA; embolus very long, originating prolaterally, curving in broad arc distally and ending midway along the retrolateral margin of the tegulum (Haddad & Mbo 2015: figs 37, 38, 43, 44).................................................................................................. M. megae Haddad & Mbo, 2015 - Palpal tibia with more than one apophysis; embolus shorter.................................................... 3 3. Palpal tibia bearing four retrolateral apophyses, dorsal RTA bifid; embolus short, needle-shaped, obliquely situated beneath sclerotised tegular apophysis (Haddad 2005: figs 4–6); large spiders,> 11.7 mm in length........ M. natalis (Pocock, 1898) - Palpal tibia bearing two or three retrolateral apophyses, dorsal RTA not bifid; embolus not hidden beneath tegular apophysis, slightly curved and flattened; medium to large spiders, <10 mm in length......................................... 4 4. Palpal tibia with two apophyses; dorsal RTA massive, sickle-shaped in retrolateral view, as long as dorsal surface of tibia and approximately half the length of the cymbium (Figs 17, 39).................................................... 5 - Palpal tibia with three apophyses; dorsal RTA shorter, triangular (Haddad & Mbo 2015: fig. 54) or hatchet-shaped (Haddad 2013: fig. 38), clearly shorter than dorsal surface of tibia and less than half the length of the cymbium.................. 6 5. Tegulum with distinct spike-like prodistal tegular apophysis; embolus originating mediodistally, axis directed approximately towards 1 o’clock (Figs 32, 37).................................................. M. tigris Haddad & Mbo, 2015 - Tegulum without prodistal tegular apophysis; embolus originating prodistally, broad and flattened, slightly curved and transversely oriented (Figs 16, 24)...................................................... M. mygaloides sp. nov. 6. Dorsal RTA triangular, originating dorsally above two smaller teeth-like ventral apophyses (Haddad & Mbo 2015: figs 39, 40, 53, 54).................................................................... M. tropicus Haddad & Mbo, 2015 - Dorsal RTA stout, hatchet-shaped, slightly curved ventrally in retrolateral view, originating dorsally above tongue-like ventral RTA, with small lobed apophysis between them proximally (Haddad 2013: figs 36, 37)............ M. martini Simon, 1898 7. Epigyne with horseshoe-shaped ridges; cheliceral promarginal pronounced mound absent............................ 8 - Epigyne with oval- or slit-shaped ridges; cheliceral promarginal pronounced mound present.......................... 9 8. Lateral margins of epigynal ridges almost parallel but converging slightly posteriorly, forming subrectangular atrium (Haddad & Mbo 2015: figs 33, 45); copulatory ducts longitudinal to epigynal axis; ST II without anterior pore-bearing receptacle (Haddad & Mbo 2015: fig. 46)............................................. M. meridionalis Haddad & Mbo, 2015 - Lateral margins of epigynal ridges broadening medially, forming diamond-shaped atrium (Haddad 2013: fig. 34); copulatory openings obliquely oriented to epigyne axis; ST II with anterior pore-bearing receptacle (Haddad 2013: fig. 35)............................................................................................ M. martini Simon, 1898 9. Epigyne ventrally with strongly sclerotized ridges forming complete ovals, directed anterolaterally along their axis....... 10 - Epigyne ventrally with ridges forming incomplete ovals or with curved ridges.................................... 11 10. Epigynal ridges touching medially (Haddad 2005: figs 1, 2); spermathecae linked by broad median U-shaped tube (Haddad 2005: fig. 3); large spiders,> 15 mm in length........................................... M. natalis (Pocock, 1898) - Epigynal ridges clearly separated (Haddad & Mbo 2015: fig. 41); spermathecae not connected by broad median U-shaped tube, but projecting anterolaterally (Haddad & Mbo 2015: fig. 42); small spiders, ca. 7 mm in length.................................................................................................. M. megae Haddad & Mbo, 2015 11. Epigyne with two widely separated oblique incomplete oval ridges, with their anterior margins directed anteromesally; ST II large, suboval, situated laterally (Figs 12, 13; Haddad & Mbo 2015: figs 35, 49, 50)............................... 12 - Epigyne with curved epigynal ridges, oriented more or less parallel to body axis; ST II either small, round and medially situated (Haddad & Mbo 2015: fig. 48) or with large lateral extensions projecting posteriorly (Haddad & Mbo 2015: fig. 52)...... 13 12. Epigynal ridges with distinct oval lateral lobe positioned posterior to lateral ridge (Figs 10, 12)..... M. mygaloides sp. nov. - Epigynal ridges with narrow fringe along their anterior margin, without oval lateral lobe (Fig. 46; Haddad & Mbo 2015: fig. 49)......................................................................... M. tigris Haddad & Mbo, 2015 13. Epigyne with small ridges, curved inwards, copulatory openings anteriorly in ridges (Haddad & Mbo 2015: figs 34, 47)............................................................................. M. seiugatus Haddad & Mbo, 2015 - Epigyne with large ridges, curved outwards, forming ovoid median atrium, with copulatory openings posteriorly in ridges (Haddad & Mbo 2015: figs 36, 51)............................................. M. tropicus Haddad & Mbo, 2015

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Zenodo (CERN European Organization for Nuclear Research)
Published
2025-05-21
DOI
https://doi.org/10.5281/zenodo.15595295
Primary Topic
Subterranean biodiversity and taxonomy
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article
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article

Messapus Simon 1898

Zenodo (CERN European Organization for Nuclear Research)
Subterranean biodiversity and taxonomy
article

Messapus Simon 1898

article en

Abstract

Updated key to the species of Messapus 1. Males (males of M. meridionalis Haddad & Mbo, 2015 and M. seiugatus Haddad & Mbo, 2015 unknown).............. 2 - Females............................................................................................. 7 2. Palp with single elongate slender RTA; embolus very long, originating prolaterally, curving in broad arc distally and ending midway along the retrolateral margin of the tegulum (Haddad & Mbo 2015: figs 37, 38, 43, 44).................................................................................................. M. megae Haddad & Mbo, 2015 - Palpal tibia with more than one apophysis; embolus shorter.................................................... 3 3. Palpal tibia bearing four retrolateral apophyses, dorsal RTA bifid; embolus short, needle-shaped, obliquely situated beneath sclerotised tegular apophysis (Haddad 2005: figs 4–6); large spiders,> 11.7 mm in length........ M. natalis (Pocock, 1898) - Palpal tibia bearing two or three retrolateral apophyses, dorsal RTA not bifid; embolus not hidden beneath tegular apophysis, slightly curved and flattened; medium to large spiders, <10 mm in length......................................... 4 4. Palpal tibia with two apophyses; dorsal RTA massive, sickle-shaped in retrolateral view, as long as dorsal surface of tibia and approximately half the length of the cymbium (Figs 17, 39).................................................... 5 - Palpal tibia with three apophyses; dorsal RTA shorter, triangular (Haddad & Mbo 2015: fig. 54) or hatchet-shaped (Haddad 2013: fig. 38), clearly shorter than dorsal surface of tibia and less than half the length of the cymbium.................. 6 5. Tegulum with distinct spike-like prodistal tegular apophysis; embolus originating mediodistally, axis directed approximately towards 1 o’clock (Figs 32, 37).................................................. M. tigris Haddad & Mbo, 2015 - Tegulum without prodistal tegular apophysis; embolus originating prodistally, broad and flattened, slightly curved and transversely oriented (Figs 16, 24)...................................................... M. mygaloides sp. nov. 6. Dorsal RTA triangular, originating dorsally above two smaller teeth-like ventral apophyses (Haddad & Mbo 2015: figs 39, 40, 53, 54).................................................................... M. tropicus Haddad & Mbo, 2015 - Dorsal RTA stout, hatchet-shaped, slightly curved ventrally in retrolateral view, originating dorsally above tongue-like ventral RTA, with small lobed apophysis between them proximally (Haddad 2013: figs 36, 37)............ M. martini Simon, 1898 7. Epigyne with horseshoe-shaped ridges; cheliceral promarginal pronounced mound absent............................ 8 - Epigyne with oval- or slit-shaped ridges; cheliceral promarginal pronounced mound present.......................... 9 8. Lateral margins of epigynal ridges almost parallel but converging slightly posteriorly, forming subrectangular atrium (Haddad & Mbo 2015: figs 33, 45); copulatory ducts longitudinal to epigynal axis; ST II without anterior pore-bearing receptacle (Haddad & Mbo 2015: fig. 46)............................................. M. meridionalis Haddad & Mbo, 2015 - Lateral margins of epigynal ridges broadening medially, forming diamond-shaped atrium (Haddad 2013: fig. 34); copulatory openings obliquely oriented to epigyne axis; ST II with anterior pore-bearing receptacle (Haddad 2013: fig. 35)............................................................................................ M. martini Simon, 1898 9. Epigyne ventrally with strongly sclerotized ridges forming complete ovals, directed anterolaterally along their axis....... 10 - Epigyne ventrally with ridges forming incomplete ovals or with curved ridges.................................... 11 10. Epigynal ridges touching medially (Haddad 2005: figs 1, 2); spermathecae linked by broad median U-shaped tube (Haddad 2005: fig. 3); large spiders,> 15 mm in length........................................... M. natalis (Pocock, 1898) - Epigynal ridges clearly separated (Haddad & Mbo 2015: fig. 41); spermathecae not connected by broad median U-shaped tube, but projecting anterolaterally (Haddad & Mbo 2015: fig. 42); small spiders, ca. 7 mm in length.................................................................................................. M. megae Haddad & Mbo, 2015 11. Epigyne with two widely separated oblique incomplete oval ridges, with their anterior margins directed anteromesally; ST II large, suboval, situated laterally (Figs 12, 13; Haddad & Mbo 2015: figs 35, 49, 50)............................... 12 - Epigyne with curved epigynal ridges, oriented more or less parallel to body axis; ST II either small, round and medially situated (Haddad & Mbo 2015: fig. 48) or with large lateral extensions projecting posteriorly (Haddad & Mbo 2015: fig. 52)...... 13 12. Epigynal ridges with distinct oval lateral lobe positioned posterior to lateral ridge (Figs 10, 12)..... M. mygaloides sp. nov. - Epigynal ridges with narrow fringe along their anterior margin, without oval lateral lobe (Fig. 46; Haddad & Mbo 2015: fig. 49)......................................................................... M. tigris Haddad & Mbo, 2015 13. Epigyne with small ridges, curved inwards, copulatory openings anteriorly in ridges (Haddad & Mbo 2015: figs 34, 47)............................................................................. M. seiugatus Haddad & Mbo, 2015 - Epigyne with large ridges, curved outwards, forming ovoid median atrium, with copulatory openings posteriorly in ridges (Haddad & Mbo 2015: figs 36, 51)............................................. M. tropicus Haddad & Mbo, 2015

Zenodo (CERN European Organization for Nuclear Research)
University of the Free State (ZA), Bernardino Rivadavia Natural Sciences Museum (AR)
Openalex Percentile: Top 76%
Subterranean biodiversity and taxonomy
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