Collitoptyx (Insolitoptyx) Volynkin & Černý, 2024, subgen. n.

Subgenus Insolitoptyx subgen. n. https://zoobank.org/ urn:lsid:zoobank.org:act: D6CD19EF-443F-4ED2-85EF-B2E25C8F03FE Type species: Collitoptyx (Insolitoptyx) meghalaya sp. n., by present designation. Diagnosis. In the male genitalia, the new subgenus (Figs 42, 43) differs from the nominate one (Figs 34–41) in the following characters. (1) The vinculum is longer and has dilated sclerotised ventral ends of the arms and thinner ‘bridge’ between them. (2) The dorsal sclerotised triangular part of the juxta is very short. (3) The anterior (cranial) side of the juxtal fold is membranous and lacking the caulis. (4) The juxta is connected with the sacculi with a considerably shorter membrane. (5) The costa is shorter, occupying the proximal third of the dorsal margin of the valva while the valvula is markedly longer and broader. (6) The conjuga is short and rectangular, with weakly sclerotised apex, separated from its counterpart by the membrane of diaphragm. (7) The lamella centralis is considerably longer and distally downcurved. (8) The sacculus is narrower, distally tapered and lacks the posterior-ventral process. (9) The distal saccular process is bilobate, its distal lobe is short but broad, triangular and spinulose while the dorsal lobe is ca. 3 times longer than the distal one, thorn-shaped and directed dorsad. This combination of two lobes with a triangular and dentate distal one makes an illusion that the latter is homologous to the dorsal saccular process of Collitoptyx s. str., which is displaced more distally, but in Collitoptyx (Insolitoptyx) subgen. n. the clasper region is situated at the base of this triangular process proving that it is homologous to the distal saccular process of Collitoptyx s. str. having a clasper region at its base. (10) The anellus bears two large ventro-lateral sclerotised plates. (11) The main diverticulum of the vesica has a utricular ventral subdiverticulum, which is distally bilobate with the proximal (and shorter) lobe bearing a tiny conical cornutus, whereas in the nominate subgenus the cornutus is situated on the wall of the main diverticulum. The female genitalia of Collitoptyx (Insolitoptyx) subgen. n. (Fig. 58) differ clearly from the nominate subgenus (Figs 52–57) in the longer ductus bursae having a sclerotised anterior section, and the pyriform corpus bursae with a short, narrow and membranous posterior section whereas in Collitoptyx (Collitoptyx) the posterior section is long, broad and with a large ventral or lateral sclerotised protrusion. Etymology. The genus name is an aggregate of the Latin adjective ‘insolitus’ meaning ‘unusual’ and the genus-group name Thysanoptyx. The name refers to the unusual structure of the distal saccular process of the type species. Gender is feminine.