Cranionycta (Nefrendinycta) senjelungma Kiss, 2025, sp. nov.

Cranionycta (Nefrendinycta) senjelungma sp. nov. urn:lsid:zoobank.org:act: E67CE664-5FD4-4C17-9F26-C93748B190C5 (Figs 33, 34, 70, 102, 103, 122, 141) Holotype. Female, Nepal, Kangchenjunga, Deorali Danda, Anpan, 1900 m, 6–7.viii.2000, leg. M. Hreblay & T. Csővári, slide No.: KA2528f (PG). Paratypes. Nepal. 1 male, same data as holotype, slide No.: KA 963m (PG); 1 female, Kangchenjunga, Milke Danda, Nesum, 1500 m, 2–3.viii.2000, leg. M. Hreblay & T. Csővári, slide No.: KA962f (PG). Diagnosis. Cranionycta (Nefrendinycta) senjelungma (Figs 33, 34) is externally similar to C. (N.) warreni (Figs 31, 21), C. (N.) dierli (Fig. 35) and especially to C. (N.) mailungana (Fig. 26), C. (N.) keeskleini (Figs 39, 40), C. (N.) punakhae (Fig. 42) and the male of C. (N.) fuscothoracica (Fig. 44). It can be distinguished from all its relatives by the narrower costal area with ochreous shading, and the wider inner part of the medial field at the basal area; from C. (N.) warreni by its generally larger size (wingspan 30–31 mm vs. 28–29 mm of C. (N.) warreni); from C. (N.) warreni and C. (N.) dierli by the generally more warmly coloured forewing with more ochreous shading and more contrasting dark wing pattern elements; the more ochreous basal area; the more contrastingly marked antemedial line; and the larger, obscure orbicular stigma with a rather conspicuous blackish line inwardly. It can be distinguished from C. (N.) mailungana, C. (N.) keeskleini, C. (N.) punakhae and the male of C. (N.) fuscothoracica by the somewhat more homogenous, less mottled forewing colouration; the larger, more homogenously, ochreousfilled basal area; and the more pronounced blackish curve as a remnant of the inner outline of the orbicular stigma; from C. (N.) keeskleini by the uniformly more brownish hindwing with a more or less absent postdiscal line. The male genitalia (Fig. 70) can be separated from those of C. (N.) warreni (Fig. 69) by the somewhat slenderer uncus; and the somewhat narrower, longer, and slightly more parallel valvae with a more gently curved ventral margin apically, and a narrower apex. The female genitalia (Figs 102, 103) differ from those of C. (N.) dierli (Fig. 101) by the somewhat shorter ductus bursae; the relatively narrower, longer junction of the appendix bursae with the corpus bursae; the basally somewhat straighter appendix bursae with a shorter basal curve of the sclerotized section; the less angled, straighter, somewhat longer terminal twist of the appendix bursae; and the longer appendix bursae relative to the corpus bursae; from those of C. (N.) warreni (Fig. 100) by the somewhat longer, narrower ductus bursae; the wider, somewhat shorter junction of the appendix bursae with the corpus bursae; and the gently curved appendix bursae with a larger basal curve of the sclerotized section, a gently curved, laterally oriented sclerotization, and an obliquely positioned, roughly narrowing terminal twist. In the male terminal abdominal segments (Fig. 122), compared to C. (N.) warreni (Fig. 121), C. (N.) senjelungma has a shorter 8 th sternite with a more irregular, indistinct, cloverleaf-shaped sclerotized distal patch, with a narrower and constricted sclerotized stripe in the lower window; a slightly narrower 8 th tergite with less pointed proximal edge, a smaller distal end of lateral sides, and a proportionally narrower window. The anterolateral sclerites of the 8 th abdominal segment are shorter than those of C. (N.) warreni. In the female 7 th abdominal segments (Fig. 141), the sternite has wavier lateral sides with slightly rounded distal corner than C. (N.) warreni (Fig. 139) and C. (N.) dierli (Fig. 140); a somewhat narrower, slightly outstanding sclerotized semi-circular distal band than C. (N.) dierli. Description. Imago (Figs 33, 34). Wingspan 30–31 mm. Head. Relatively large; 2 nd segment of palpus longer than 3 rd in both sexes; antennae of both sexes filiform. Thorax. Moderately wide, greyish-white with two blackish spots; patagia concolorous with thorax with thin blackish outline; tegulae concolorous with thorax with thin blackish outline. Abdomen. Whitish-grey with some darker middorsal scale tufts. Wing. Forewing narrow, apically somewhat elongate and obtuse, dark greyish-brown with ochreous shading in basal, costal and marginal areas; basal spot blackish, small, more or less semi-circular; basal streak blackish, short, wedge-shaped; tornal streak blackish, thin, long, indistinct between medial and postmedial lines, widest between postmedial and terminal lines; apical dash tiny, indistinct blackish patch; basal line blackish, double, outer line conspicuously wider than inner line, inner line very short, dot-like, filled with greyish-brown; antemedial line reduced and interrupted, with three blackish spots at costal area, and filled with whitish at inner margin with indistinct outward blackish spot; medial line blackish, wide, zigzag; medial fascia relatively narrow, blackish, outwardly fading; postmedial line double, crenulate, inner line greyish-black, thin, reduced, outer line blackish, more prominent, with narrow black stripe between veins R3 and M2, filled with whitish; subterminal line off-white, narrow, indistinct; terminal line whitish, interrupted by black on veins; basal field greyish-brown, narrow, fused with suprabasal patch, antemedial field (in male) and interior of basal line, and joined to subbasal patch; subbasal patch whitish with ochreous shading; suprabasal patch brownish-ochre, obscure, stripe-like; antemedial field greyish-brown with ochreous shading and blackish patch at costal area; medial field inwardly whitish with ochreous shading at costal area, outwardly greyish-brown, inner part narrow but basally widened, outer part narrow; subterminal field greyish-brown with some ochreous shading and large, blackish-grey, squarish patches in costal area, obscure between veins M1 and M2 and around tornal streak; terminal field greyish-black, narrow, indistinct; orbicular stigma reduced, whitish with short, black curved line or tiny dots inwardly; reniform stigma large, greyish-brown, darker inwardly, outlined with series of blackish dots and curved line; four quadrangular whitish spots at costa between medial fascia and terminal line, gradually decreasing in size towards apex; tornal patch blackish, small; fringe whitish with greyish patches. Hindwing rounded, apically slightly pointed, greyish-brown; marginal band slightly darker greyish-brown; postdiscal line and discal spot faint; tornal patch faint, slightly darker greyish-ochreous; fringe whitish with greyish patches. Male genitalia (Fig. 70). Uncus moderately short, sclerotized, hairy, narrowing distally; apex hooked. Scaphium moderately sclerotized, pliers-like; subscaphium weakly sclerotized. Tegumen long with lobe-like peniculus covered with dense long hairs. Saccus sclerotized, V-shaped. Juxta simple, moderately sclerotized, widest at base, shield-shaped. Manica double, moderately sclerotized lobe-like extension with spinulose structure supporting numerous small spinules. Transtilla short, rod-like, uniformly wide, sclerotized. Valvae elongate, narrowest in basal third then widening stepwise towards apex, with two breaks on costa; sacculus narrow, sclerotized; clavus slightly curved; valval androconial apparatus absent; medial sclerite straight and sclerotized; apex gently curved ventrally and with wide, dense corona. Phallus simple, tubular, moderately sclerotized; carina slightly more sclerotized, wide, short, wedge-shaped. Vesica as long as phallus, moderately wide, essentially tubular with three basal diverticula one of them bearing spinulose patch, distal diverticulum larger, elongate; tooth-like cornuti absent from mid-lateral surface. Female genitalia (Figs 102, 103). Ovipositor rounded, slightly longer than wide, tapering posteriorly. Papillae anales weakly sclerotized, oval, densely hairy. Anterior and posterior apophysis equal in length, weakly sclerotized, rod-like. Ostium longitudinally ribbed, moderately sclerotized. Antrum short, moderately sclerotized, more or less funnel-shaped. Ductus bursae short, moderately sclerotized, ribbed. Corpus bursae large, bulbous, weakly sclerotized with two opposing distal signa. Appendix bursae shorter than corpus bursae, coiled, moderately sclerotized, ribbed except in distal third, terminal twist short and straight; junction with corpus bursae long and narrow. Male 7 th and 8 th abdominal segments (Fig. 122). 7 th sternite roughly trapezoidal, wider than long, uniformly sclerotized with slightly more sclerotized, narrow semi-circular distal band; proximal edges slightly wavy, laterally extended; lateral sides slightly concave with less contrasting margin; distal edge slightly concave. 7 th tergite quadrangular, wider than long; proximally slightly sclerotized, distally with slightly more sclerotized semi-circular band, and with indistinct window in middle; proximal edge more or less straight with two long, curved, slightly stronger sclerotized rods; lateral sides proximally concave then distally slightly convex with less contrasting margin; distal edge slightly concave. 8 th sternite roughly trapezoidal with rounded corner, as long as wide; window relatively large, more or less quadrangular with rounded proximal margin and angled distal margin, with narrow, slightly constricted, pointed sclerotized stripe proximally; proximal edge slightly arched with sclerotized tiny patch in middle; lateral side proximally uniformly narrow, distally strongly widened and fused with irregular and indistinct cloverleaf-shaped sclerotized patch; distal edge concave. 8 th tergite sclerotized proximally, bell-shaped, about 1.5× as long as wide; window spade-shaped, long, with more or less contrast margin; proximal edge short, pointed; lateral sides proximally wider, narrowest in middle, and with wide, rounded spur-like distal end; distal edge more or less straight. Anterolateral sclerites of 8 th segment

Publication Details

Journal
Zenodo (CERN European Organization for Nuclear Research)
Published
2025-05-26
DOI
https://doi.org/10.5281/zenodo.15563231
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Aquatic Ecosystems and Phytoplankton Dynamics
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article

Cranionycta (Nefrendinycta) senjelungma Kiss, 2025, sp. nov.

Zenodo (CERN European Organization for Nuclear Research)
Aquatic Ecosystems and Phytoplankton Dynamics
article

Cranionycta (Nefrendinycta) senjelungma Kiss, 2025, sp. nov.

article en

Abstract

Cranionycta (Nefrendinycta) senjelungma sp. nov. urn:lsid:zoobank.org:act: E67CE664-5FD4-4C17-9F26-C93748B190C5 (Figs 33, 34, 70, 102, 103, 122, 141) Holotype. Female, Nepal, Kangchenjunga, Deorali Danda, Anpan, 1900 m, 6–7.viii.2000, leg. M. Hreblay & T. Csővári, slide No.: KA2528f (PG). Paratypes. Nepal. 1 male, same data as holotype, slide No.: KA 963m (PG); 1 female, Kangchenjunga, Milke Danda, Nesum, 1500 m, 2–3.viii.2000, leg. M. Hreblay & T. Csővári, slide No.: KA962f (PG). Diagnosis. Cranionycta (Nefrendinycta) senjelungma (Figs 33, 34) is externally similar to C. (N.) warreni (Figs 31, 21), C. (N.) dierli (Fig. 35) and especially to C. (N.) mailungana (Fig. 26), C. (N.) keeskleini (Figs 39, 40), C. (N.) punakhae (Fig. 42) and the male of C. (N.) fuscothoracica (Fig. 44). It can be distinguished from all its relatives by the narrower costal area with ochreous shading, and the wider inner part of the medial field at the basal area; from C. (N.) warreni by its generally larger size (wingspan 30–31 mm vs. 28–29 mm of C. (N.) warreni); from C. (N.) warreni and C. (N.) dierli by the generally more warmly coloured forewing with more ochreous shading and more contrasting dark wing pattern elements; the more ochreous basal area; the more contrastingly marked antemedial line; and the larger, obscure orbicular stigma with a rather conspicuous blackish line inwardly. It can be distinguished from C. (N.) mailungana, C. (N.) keeskleini, C. (N.) punakhae and the male of C. (N.) fuscothoracica by the somewhat more homogenous, less mottled forewing colouration; the larger, more homogenously, ochreousfilled basal area; and the more pronounced blackish curve as a remnant of the inner outline of the orbicular stigma; from C. (N.) keeskleini by the uniformly more brownish hindwing with a more or less absent postdiscal line. The male genitalia (Fig. 70) can be separated from those of C. (N.) warreni (Fig. 69) by the somewhat slenderer uncus; and the somewhat narrower, longer, and slightly more parallel valvae with a more gently curved ventral margin apically, and a narrower apex. The female genitalia (Figs 102, 103) differ from those of C. (N.) dierli (Fig. 101) by the somewhat shorter ductus bursae; the relatively narrower, longer junction of the appendix bursae with the corpus bursae; the basally somewhat straighter appendix bursae with a shorter basal curve of the sclerotized section; the less angled, straighter, somewhat longer terminal twist of the appendix bursae; and the longer appendix bursae relative to the corpus bursae; from those of C. (N.) warreni (Fig. 100) by the somewhat longer, narrower ductus bursae; the wider, somewhat shorter junction of the appendix bursae with the corpus bursae; and the gently curved appendix bursae with a larger basal curve of the sclerotized section, a gently curved, laterally oriented sclerotization, and an obliquely positioned, roughly narrowing terminal twist. In the male terminal abdominal segments (Fig. 122), compared to C. (N.) warreni (Fig. 121), C. (N.) senjelungma has a shorter 8 th sternite with a more irregular, indistinct, cloverleaf-shaped sclerotized distal patch, with a narrower and constricted sclerotized stripe in the lower window; a slightly narrower 8 th tergite with less pointed proximal edge, a smaller distal end of lateral sides, and a proportionally narrower window. The anterolateral sclerites of the 8 th abdominal segment are shorter than those of C. (N.) warreni. In the female 7 th abdominal segments (Fig. 141), the sternite has wavier lateral sides with slightly rounded distal corner than C. (N.) warreni (Fig. 139) and C. (N.) dierli (Fig. 140); a somewhat narrower, slightly outstanding sclerotized semi-circular distal band than C. (N.) dierli. Description. Imago (Figs 33, 34). Wingspan 30–31 mm. Head. Relatively large; 2 nd segment of palpus longer than 3 rd in both sexes; antennae of both sexes filiform. Thorax. Moderately wide, greyish-white with two blackish spots; patagia concolorous with thorax with thin blackish outline; tegulae concolorous with thorax with thin blackish outline. Abdomen. Whitish-grey with some darker middorsal scale tufts. Wing. Forewing narrow, apically somewhat elongate and obtuse, dark greyish-brown with ochreous shading in basal, costal and marginal areas; basal spot blackish, small, more or less semi-circular; basal streak blackish, short, wedge-shaped; tornal streak blackish, thin, long, indistinct between medial and postmedial lines, widest between postmedial and terminal lines; apical dash tiny, indistinct blackish patch; basal line blackish, double, outer line conspicuously wider than inner line, inner line very short, dot-like, filled with greyish-brown; antemedial line reduced and interrupted, with three blackish spots at costal area, and filled with whitish at inner margin with indistinct outward blackish spot; medial line blackish, wide, zigzag; medial fascia relatively narrow, blackish, outwardly fading; postmedial line double, crenulate, inner line greyish-black, thin, reduced, outer line blackish, more prominent, with narrow black stripe between veins R3 and M2, filled with whitish; subterminal line off-white, narrow, indistinct; terminal line whitish, interrupted by black on veins; basal field greyish-brown, narrow, fused with suprabasal patch, antemedial field (in male) and interior of basal line, and joined to subbasal patch; subbasal patch whitish with ochreous shading; suprabasal patch brownish-ochre, obscure, stripe-like; antemedial field greyish-brown with ochreous shading and blackish patch at costal area; medial field inwardly whitish with ochreous shading at costal area, outwardly greyish-brown, inner part narrow but basally widened, outer part narrow; subterminal field greyish-brown with some ochreous shading and large, blackish-grey, squarish patches in costal area, obscure between veins M1 and M2 and around tornal streak; terminal field greyish-black, narrow, indistinct; orbicular stigma reduced, whitish with short, black curved line or tiny dots inwardly; reniform stigma large, greyish-brown, darker inwardly, outlined with series of blackish dots and curved line; four quadrangular whitish spots at costa between medial fascia and terminal line, gradually decreasing in size towards apex; tornal patch blackish, small; fringe whitish with greyish patches. Hindwing rounded, apically slightly pointed, greyish-brown; marginal band slightly darker greyish-brown; postdiscal line and discal spot faint; tornal patch faint, slightly darker greyish-ochreous; fringe whitish with greyish patches. Male genitalia (Fig. 70). Uncus moderately short, sclerotized, hairy, narrowing distally; apex hooked. Scaphium moderately sclerotized, pliers-like; subscaphium weakly sclerotized. Tegumen long with lobe-like peniculus covered with dense long hairs. Saccus sclerotized, V-shaped. Juxta simple, moderately sclerotized, widest at base, shield-shaped. Manica double, moderately sclerotized lobe-like extension with spinulose structure supporting numerous small spinules. Transtilla short, rod-like, uniformly wide, sclerotized. Valvae elongate, narrowest in basal third then widening stepwise towards apex, with two breaks on costa; sacculus narrow, sclerotized; clavus slightly curved; valval androconial apparatus absent; medial sclerite straight and sclerotized; apex gently curved ventrally and with wide, dense corona. Phallus simple, tubular, moderately sclerotized; carina slightly more sclerotized, wide, short, wedge-shaped. Vesica as long as phallus, moderately wide, essentially tubular with three basal diverticula one of them bearing spinulose patch, distal diverticulum larger, elongate; tooth-like cornuti absent from mid-lateral surface. Female genitalia (Figs 102, 103). Ovipositor rounded, slightly longer than wide, tapering posteriorly. Papillae anales weakly sclerotized, oval, densely hairy. Anterior and posterior apophysis equal in length, weakly sclerotized, rod-like. Ostium longitudinally ribbed, moderately sclerotized. Antrum short, moderately sclerotized, more or less funnel-shaped. Ductus bursae short, moderately sclerotized, ribbed. Corpus bursae large, bulbous, weakly sclerotized with two opposing distal signa. Appendix bursae shorter than corpus bursae, coiled, moderately sclerotized, ribbed except in distal third, terminal twist short and straight; junction with corpus bursae long and narrow. Male 7 th and 8 th abdominal segments (Fig. 122). 7 th sternite roughly trapezoidal, wider than long, uniformly sclerotized with slightly more sclerotized, narrow semi-circular distal band; proximal edges slightly wavy, laterally extended; lateral sides slightly concave with less contrasting margin; distal edge slightly concave. 7 th tergite quadrangular, wider than long; proximally slightly sclerotized, distally with slightly more sclerotized semi-circular band, and with indistinct window in middle; proximal edge more or less straight with two long, curved, slightly stronger sclerotized rods; lateral sides proximally concave then distally slightly convex with less contrasting margin; distal edge slightly concave. 8 th sternite roughly trapezoidal with rounded corner, as long as wide; window relatively large, more or less quadrangular with rounded proximal margin and angled distal margin, with narrow, slightly constricted, pointed sclerotized stripe proximally; proximal edge slightly arched with sclerotized tiny patch in middle; lateral side proximally uniformly narrow, distally strongly widened and fused with irregular and indistinct cloverleaf-shaped sclerotized patch; distal edge concave. 8 th tergite sclerotized proximally, bell-shaped, about 1.5× as long as wide; window spade-shaped, long, with more or less contrast margin; proximal edge short, pointed; lateral sides proximally wider, narrowest in middle, and with wide, rounded spur-like distal end; distal edge more or less straight. Anterolateral sclerites of 8 th segment

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