Inella harryleei Rolan & Fernandez-Garces 2008

“ Inella ” harryleei Rolán & Fernández-Garcés, 2008 Fig. 1 Inella harryleei Rolán & Fernández-Garcés, 2008: 105, fig. 13a–k. “ Inella ” harryleei – Fernandes & Pimenta 2019a: fig. 3k–l; 2019b: 46, figs 2l, 30–32. Inella harryleei – Leal 2021: 6, fig. 38. Type material Holotype USA • sh; Florida, off Dry Tortugas; depth 90 m; FLMNH 419182. Paratypes See Rolán & Fernández-Garcés (2008). Material examined GUADELOUPE (Karubenthos 2 expedition) • 2 sh; stn DW4592; MNHN. Emended description Shell sinistral, conical-fusiform, up to 15.9 mm long, 2.4 mm wide, length/width ratio 6.6, apical angle of early whorls 11°. Protoconch paucispiral, three whorls, 0.96 mm long, 0.57 mm wide; first whorl smooth, inflated, only slightly narrower than subsequent whorls, which have two thin, nearly smooth spiral cords, situated at 33% and 76% of last whorl height; transition to teleoconch nearly indistinct. Teleoconch with up to 20 whorls; two spiral cords (adapical and abapical) in beginning of teleoconch, continuous to those of protoconch, but abapical one considerably more developed until body whorl; median spiral cord emerges very narrowly at end of second whorl, slowly developing but never reaching same size as other cords; suture shallow, with a smooth to slightly wavy sutural cord; 16–17 opisthocline axial ribs on 12 th teleoconch whorl; medium to large-sized (on abapical cord), rounded to slightly elliptical nodules; nodulous, moderately thin subperipheral cord, with a slightly nodulous adapical basal cord right below it and a thin, nearly smooth abapical basal cord situated apart; no evident supranumerical cord; nearly rounded aperture, 1.1 mm long, 0.84 mm wide, length/width ratio 1.3; anterior canal very long, partially open, directed downward, 1.0 mm long, 0.45 mm wide, length/width ratio 2.3. Shell mainly white, stained by few and discrete light brown (or cream) axial patches, usually comprising width of one or two axial ribs. Remarks The only known protoconch of “ I.” harryleei from Guadeloupe (Fig. 1C) has a first whorl apparently more inflated than those illustrated in the original description, from Florida and Louisiana (USA), although all other shell features are very similar. Based on the nearest record from the Florida Keys (Rolán & Fernández-Garcés 2008), the present record extends the known range of “ I.” harryleei by ~ 2180 km into the Caribbean. However, Rolán & Fernández-Garcés (2008) indicated a shell fragment from the ‘West Indies’ as possibly belonging to this species. Lamy & Pointier (2018: 284, pl. 91 fig. 7a–b) identified a shell from Guadeloupe as Inella aff. harryleei, following the morph from Florida illustrated by Rolán & Fernández-Garcés (2008) under this name. In fact, the teleoconch of this shell from Guadeloupe differs from that of “ I.” harryleei (see Rolán & Fernández-Garcés 2008 for further details). Moreover, this shell has an abapical spiral cord that is even more prominent than that of “ I.” aff. harryleei from Florida. Unfortunately, the broken apex of the shell from Guadeloupe precluded further comparisons. Lamy & Pointier (2018) indicated a depth range of 55–360 m for this morph; considering that “ I.” aff. harryleei from Florida is only known from 55 m (Rolán & Fernández-Garcés 2008), only the depth of 360 m is confidently assigned to this peculiar shell from Guadeloupe. Geographic distribution USA: Florida (Rolán & Fernández-Garcés 2008; Fernandes & Pimenta 2019b), Louisiana (Rolán & Fernández-Garcés 2008; Garcia & Lee 2020); Guadeloupe (this study). Bathymetric distribution Empty shells previously known from 46–500 m (Rolán & Fernández-Garcés 2008), live specimens only known from 63 m (Fernandes & Pimenta 2019b). This study: 201–214 m (empty shells).