Spungisomyia germanica Jaschhof, 2024, sp. nov.

Spungisomyia germanica sp. nov. urn:lsid:zoobank.org:act: F02F38B6-4C58-4E06-8EB4-2DC294D33C04 Fig. 31 Diagnosis Morphology Spungisomyia germanica sp. nov. is the only species of Spungisomyia known to have two small, clearly separate prescutellar windows that are in juxtaposition with each other; the usual condition in this genus is a single, large prescutellar window. As regards male terminalia characters (Fig. 31A), the compact gonostylus has a conspicuously thick base, which in ventral view is indicated by the strongly convex outer margin next to the basolateral apophysis (↓ 1); the small, U-shaped gonocoxal emargination has a broad, darkly pigmented margin (↓ 2); and the conspicuously thin parameres are multipointed apically (↓ 3). Larvae and females of the new species are unknown. DNA barcode CO1 sequences (651‒674bp) of the type specimens are available in BIN: BOLD:AEO6595. Search on BOLDʼs BIN Database retrieved no further result for this BIN (accessed 27 Aug. 2023). Etymology The specific name, an adjective, refers to Germany as the country of provenance of the type material. Type material Holotype GERMANY – Baden-Württemberg • ♂; Malsch, Luderbusch; 48°91′44″ N, 8°33′24″ E; elev. 112 m; 15‒29 Mar. 2020; D. Doczkal and K. Grabow leg.; Malaise trap; willow/aspen forest and game browsing field; BOLD GBDTA10043-21; ZSM-DIP-42304-F08. Paratypes GERMANY – Baden-Württemberg • 1 ♂; same data as the holotype; BOLD GBDTA10040-21; ZSM-DIP-42304-F05 • 1 ♂; same data as for preceding; BOLD GBDTA10042-21; ZSM-DIP-42304-F07 • 1 ♂; same data as for preceding; BOLD GBDTA10045-21; ZSM-DIP-42304-F10 • 1 ♂; same data as for preceding but 48°91′56″ N, 8°33′20″ E; elev. 110 m; 1‒15 Mar. 2020; willow scrub at pond edge; BOLD GBDTA10015-21; ZSM-DIP-42304-D04 • 1 ♂; same data as for preceding; BOLD GBDTA10016-21; ZSM-DIP-42304-D05 • 1 ♂; same data as for preceding but 48°91′28″ N, 8°33′35″ E; elev. 111 m; 1‒15 Mar. 2020; willow scrub and reed; BOLD DTIII8528-22; ZSM-DIP-42474-G01 • 1 ♂; same data as for preceding; BOLD DTIII8529-22; ZSM-DIP-42474-G02 • 1 ♂; same data as for preceding; BOLD DTIII8532-22; ZSM-DIP-42474-G05 • 1 ♂; same data as for preceding but 15‒29 Mar. 2020; BOLD DTIII8503-22; ZSM-DIP-42474-D12 • 1 ♂; same data as for preceding; BOLD DTIII8504-22; ZSM-DIP-42474-E01 • 1 ♂; same data as for preceding; BOLD DTIII8505-22; ZSM-DIP-42474-E02 • 1 ♂; same data as for preceding; BOLD DTIII8507-22; ZSM-DIP-42474-E04 • 1 ♂; same data as for preceding; BOLD DTIII8509-22; ZSM-DIP-42474-E06 • 1 ♂; same data as for preceding; BOLD DTIII8511-22; ZSM-DIP-42474-E08 • 1 ♂; same data as for preceding; BOLD DTIII8512-22; ZSM-DIP-42474-E09 • 1 ♂; same data as for preceding; BOLD DTIII8513-22; ZSM-DIP-42474-E10 • 1 ♂; same data as for preceding; BOLD DTIII8514-22; ZSM-DIP-42474-E11 • 1 ♂; same data as for preceding; BOLD DTIII8515-22; ZSM-DIP-42474-E12 • 1 ♂; same data as for preceding; BOLD DTIII8516-22; ZSM-DIP-42474-F01 • 1 ♂; same data as for preceding; BOLD DTIII8517-22; ZSM-DIP-42474-F02 • 1 ♂; same data as for preceding; BOLD DTIII8518-22; ZSM-DIP-42474-F03 • 1 ♂; same data as for preceding; BOLD DTIII8519-22; ZSM-DIP-42474-F04 • 1 ♂; same data as for preceding; BOLD DTIII8520-22; ZSM-DIP-42474-F05 • 1 ♂; same data as for preceding; BOLD DTIII8521-22; ZSM-DIP-42474-F06 • 1 ♂; same data as for preceding; BOLD DTIII8522-22; ZSM-DIP-42474-F07 • 1 ♂; same data as for preceding; BOLD DTIII8524-22; ZSM-DIP-42474-F09 • 1 ♂; same data as for preceding; BOLD DTIII8526-22; ZSM-DIP-42474-F11. – Bavaria • 1 ♂; Weilheim, Pähl, Hartschimmelhof; 47°93′86″ N, 11°18′62″ E; 20 Mar.‒24 Apr. 2020; GBOL leg.; Malaise trap; nutrient-poor meadow; BOLD DTIII8449-22; ZSM-DIP-42473-G10 • 1 ♂; same data as for preceding; BOLD DTIII8450-22; ZSM-DIP-42473-H05 • 1 ♂; same data as for preceding; BOLD DTIII8451-22; ZSM-DIP-42473-H06 • 1 ♂; same data as for preceding; BOLD DTIII8438-22; ZSM-DIP-42473-H07 • 1 ♂; same data as for preceding; BOLD DTIII8439-22; ZSM-DIP-42473-G06 • 1 ♂; same data as for preceding; BOLD DTIII8442-22; ZSM-DIP-42473-G07 • 1 ♂; same data as for preceding but 47°94′24″ N, 11°18′23″ E; 25 Jan.‒21Mar. 2020; fen; BOLD DTIII8457-22; ZSM-DIP-42474-A02 • 1♂; same data as for preceding; 47°93′83″ N, 11°18′39″ E; 21 Mar.‒24 Apr. 2020; nutrient-poor pasture; BOLD DTIII8488-22; ZSM-DIP-42474-C09 • 1 ♂; same data as for preceding; BOLD DTIII8494-22; ZSM-DIP-42474-D03. Other characters Male BODY LENGTH. 2.3‒2.7 mm. HEAD. Eye bridge 0‒1 ommatidium long dorsally. Antenna markedly longer than body; scape and pedicel concolorous with flagellum; circumfila present on all flagellomeres; neck of fourth flagellomere 1.8 × as long as node (Fig. 31B). Palpus 4-segmented, about as long as head height, apical segment longest of all. THORAX. Presutural windows absent. Anepisternal setae 0‒1. WING. Markedly longer than body. Length/ width ratio 2.8‒2.9. M 1+2 absent. LEGS. Foreleg with femur 0.9× length of tibia, tibia 1.3× length of T 2. Claws with 3 teeth basally, the outermost largest. Empodia broad, as long as claws. TERMINALIA (Fig. 31A). Gonocoxal synsclerite markedly narrowed towards base; narrow portion with same large setae as rest of ventral surface; anterior margin straight rather than rounded; true processes absent. Gonostylus 1.5× as long as thick, slightly tapered towards apex, with setae of greatly varying sizes; frontal portion as strongly convex as basal portion; pectinate tooth of considerable size. Parameres for most of their lengths fairly thin, only apically slightly broader; parameral apodemes large, branched. Aedeagal apodeme shorter than gonocoxae, moderately sclerotized; apex vanishing into membranous cap; ducts of accessory glands evident (not illustrated). Membranous portion of aedeagus expansive (not illustrated).