Triepeolus simplex Robertson 1903

Triepeolus simplex Robertson, 1903 Figs 10A, 12P, 18D, 31 Triepeolus simplex Robertson, 1903: 285 (♀, ♂). Triepeolus simplex – Mitchell 1962: 483 (redescription). — Webb 1980: 110 (♀) (lectotype designation by W.E. LaBerge). Proposed common name Simple triepeolus. Diagnosis Unique to T. simplex within the simplex species group is the presence of only short, straight setae (as opposed to an apical/subapical fringe of dense, long (> 1 MOD), curved setae) on the male S5 (Figs 10A, 31C). Otherwise, the following morphological features in combination tell both sexes of T. simplex apart from all other species of Triepeolus: the clypeus does not have a glabrous midline (Supp. file 2: Fig. S25C); the T1 discal patch is trapezoidal or almost semicircular (Fig. 31B); T2– T4 have apical transverse bands of pale tomentum that are clearly not as broad as the bands on T1 and do not have basomedially convergent anterolateral extensions (Fig. 31A–C); and the T2–T3 apical transverse bands are yellow and somewhat removed from the apical margins of the terga whereas the T4 apical transverse band is off-white and on or very little removed from the apical margin of the tergum (Figs 10A, 31A–C). Males of T. simplex are very similar in overall appearance to those of T. lunatus (Say, 1824) (outside of the simplex species group), but in males of T. lunatus both S4 and S5 each have an apical/subapical fringe of dense, long (> 1 MOD), curved setae and in both sexes the clypeus has a well-defined glabrous midline that extends most if not all of its length. Etymology The specific epithet is a Latin adjective meaning ‘simple,’ but it is not clear from the original description why Robertson (1903) chose this name. Material examined Primary type material USA • ♀, lectotype; Illinois, Carlinville, Macoupin County; 14 Aug. 1896; C.A. Robertson leg.; INHS 62277. Secondary type material USA • 1 ♂, lectoallotype; Illinois, Carlinville, Macoupin County; 7 Aug. 1897; C.A. Robertson leg.; INHS 62278. DNA barcoded material Available. BIN: BOLD:AAW8070. Specimens examined and sequenced: CANADA • 1 ♂; Ontario, Queen’s University Biological Station; 3 Aug. 2005; Forney and Gravel leg.; BOLD sample ID: QUBS0291G11; PCYU. USA – Michigan • 1 ♀; Flynn Township Nature Center, Sanilac County; 43.2670° N, 82.9480° W; 4 Aug. 2018; T.J. Wood leg.; BOLD sample ID: CMNTO_158; WRME JBWM0398680. – North Carolina • 1 ♀; Nellie (near Cataloochee), Great Smoky Mountains National Park, Haywood County; 35.6249° N, 83.1139° W; 6 Aug. 2006; J. Gibbs and C.S. Sheffield leg.; BOLD sample ID: CMNTO_038; PCYU PYU-319. – Oklahoma • 1 ♀; Keystone Wildlife Management Area (Arkansas River), Osage County; 36.3066° N, 96.5221° W; 1 Jul. 2022; M.E. Powley leg.; BOLD sample ID: CMNTO_453; OKBS OKBS.POL.4927. – Vermont • 1 ♀; Intervale Community Farm, Burlington; 44.4955° N, 73.2072° W; 3 Sep. 2019; S. Hardy leg.; BOLD sample ID: CMNTO_097; VCE 4244. Non-barcoded material CANADA – Ontario • 1 ♀; near St-Albert; 45.2528° N, 75.2030° W; 25 Jul.–1 Aug. 2019; S. Cardinal leg.; CNC 1595850 • 1 ♀; Sixteen Mile Creek (near Hwy 407), Oakville; 19 Aug. 2004; M. Buck leg.; DEBU debu00240350 • 1 ♂; Centennial Park (Toronto); 43.6584° N, 79.5899° W; 28 Aug. 2019; S. MacKell leg.; PCYU. USA – Arkansas • 2 ♀♀; Polk County; 21 Aug. 1928; A.M. James leg.; KUNHM M.G.R. Database Nos 4615, 4616. – Georgia • 1 ♂; Morrison; USNM M.G.R. Database No. 6638 • 1 ♀; SW of Watkinsville, Oconee County; 22 Aug. 1981; C.K. Starr leg.; USNM M.G.R. Database No. 1344. – Illinois • 1 ♀; Carlinville, Macoupin County; C.A. Robertson leg.; AMNH acc 38827/M.G.R. Database No. 3612. – Indiana • 1 ♂; Conrad, Newton County; 8 Jul. 2001; R.P. Jean leg.; RJC #3059/M.G.R. Database No. 6678. – Iowa • 1 ♀; Co. 3; 19 Aug. 1933; Barker leg.; USNM M.G.R. Database No. 5895. – Kansas • 1 ♀; 5 km E of Lawrence, Douglas County; 3 Sep. 1984; R. Brooks and C. Michener leg.; KUNHM M.G.R. Database No. 4608 • 1 ♀; 5 mi W of Lawrence, Douglas County; 10 Sep. 1950; J.R. White leg.; KUNHM M.G.R. Database No. 4601 • 1 ♀; 7 km E of Lawrence, Douglas County; 15 Aug. 1990; R.L. Minckley leg.; KUNHM M.G.R. Database No. 4605 • 1 ♀; Allen County; R.H. Beamer leg.; KUNHM M.G.R. Database No. 4612 • 1 ♀; Baldwin; USNM M.G.R. Database No. 6452 • 1 ♂; same data as for preceding; Jul.; Bridwell leg.; USNM M.G.R. Database No. 6639 • 1 ♂; same data as for preceding; Jul.; J.C. Bridwell leg.; USNM M.G.R. Database No. 6640 • 2 ♂♂; DeSoto; 13 Aug. 1952; W.E. LaBerge leg.; KUNHM M.G.R. Database Nos 4580, 4588 • 1 ♀; Douglas County; F.H. Snow leg.; KUNHM M.G.R. Database No. 3939 • 1 ♀; same data as for preceding; 30 Jul. 1949; C.D. Michener leg.; KUNHM M.G.R. Database No. 4600 • 2 ♂♂; same data as for preceding; 30 Jul. 1951; C.D. Michener leg.; KUNHM M.G.R. Database Nos 4579, 4582 • 1 ♂; same data as for preceding; 2 Aug. 1951; W.E. LaBerge leg.; KUNHM M.G.R. Database No. 4585 • 1 ♀; same data as for preceding; 16 Aug. 1951; W.E. LaBerge leg.; KUNHM M.G.R. Database No. 4604 • 1 ♀; same data as for preceding; 20 Aug. 1951; W.E. LaBerge leg.; KUNHM M.G.R. Database No. 4607 • 1 ♂; same data as for preceding; 18 Aug. 1955; C.W. Rettenmeyer leg.; KUNHM M.G.R. Database No. 4589 • 1 ♂; Lawrence, Douglas County; 23 Jul. 1949; Michener and Rozen leg.; AMNH M.G.R. Database No. 3611 • 1 ♀; same data as for preceding; 9 Aug. 1950; J.R. White leg.; KUNHM M.G.R. Database No. 4598 • 2 ♂♂; same data as for preceding; 9 Aug. 1950; J.R. White leg.; KUNHM M.G.R. Database Nos 4581, 4583 • 2 ♂♂; same data as for preceding; 16 Aug. 1950; J.R. White leg.; KUNHM M.G.R. Database Nos 4584, 4587 • 1 ♀; same data as for preceding; 18 Aug. 1950; J.R. White leg.; KUNHM M.G.R. Database No. 4603 • 1 ♂; same data as for preceding; 18 Aug. 1950; J.R. White leg.; KUNHM M.G.R. Database No. 4591 • 1 ♀; same data as for preceding; 18 Aug. 1951; C.D. Michener leg.; KUNHM M.G.R. Database No. 4599 • 3 ♂♂; same data as for preceding; 11 Jun. 1954; Daly leg.; USNM M.G.R. Database Nos 1345 to 1347 • 2 ♀♀; Lawrence vicinity; 30 Jul. 1977; S. Laroca leg.; KUNHM M.G.R. Database Nos 1322, 4606 • 1 ♂; Ottawa; 20 Aug. 1950; C.D. Michener leg.; KUNHM M.G.R. Database No. 4590 • 1 ♂; Topeka; 3 Aug. 1950; C.D. Michener and J.R. White leg.; KUNHM M.G.R. Database No. 4592 • 1 ♀; University of Kansas Natural History Reservation, Douglas County; 22 Jul. 1960; KUNHM M.G.R. Database No. 4602. – Louisiana • 1 ♂; Logansport; 13 Aug. 1905; La. Crop Pest Commission leg.; USNM M.G.R. Database No. 6643. – Maryland • 1 ♀; Easton, Talbot County; 38.7683° N, 76.0754° W; 18 Jul. 2016; S.W. Droege leg.; BIML USGS-DRO 446669 • 3 ♂♂; Oxford, Talbot County; 38.6833° N, 76.1650° W; 11 Jul. 2018; S.W. Droege leg.; BIML USGS-DRO 536852, USGS-DRO 536888, USGS-DRO 536897. – Michigan • 1 ♀; Al Sabo Land Preserve (Kalamazoo); 42.2140° N, 85.6740° W; 29 Jul. 2017; T.J. Wood leg.; WRME JBWM0398679. – Minnesota • 1 ♀; Dumfries, Wabasha County; 13 Aug. 1960; J.R. Powers leg.; LACM M.G.R. Database No. 3613. – Mississippi • 1 ♂; USNM M.G.R. Database No. 6642 • 1 ♀; same data as for preceding; W.H. Ashmead leg.; USNM M.G.R. Database No. 1343 • 1 ♀; 7.3 km NE of Starkville, Oktibbeha County; 33.5100° N, 88.7369° W; 19 Jun. 2001; R. Brooks and T. Peterson leg.; KUNHM MISS1B01 002 /M.G.R. Database No. 4614 • 1 ♂; same data as for preceding; 19 Jun. 2001; R. Brooks and T. Peterson leg.; KUNHM MISS1B01 002 /M.G.R. Database No. 4597. – Missouri • 1 ♂; 12 mi E of Lebanon; 5 Aug. 1951; C.D. Michener leg.; KUNHM M.G.R. Database No. 4586 • 1 ♂; Atherton; Jul. 1954; W.M. Mann leg.; USNM M.G.R. Database No. 1349 • 1 ♂; same data as for preceding; Aug. 1954; W.M. Mann leg.; USNM M.G.R. Database No. 1348 • 1 ♀; Columbia; 5 Sep. 1967; F.D. Parker leg.; USNM M.G.R. Database No. 1342 • 1 ♀; Saline County (31 mi W of Boonville); 2 Jul. 1999; L. Packer leg.; PCYU • 1 ♂; St. Louis; 12 Aug. 1938; USNM M.G.R. Database No. 6641. – Nebraska • 1 ♀; West Point; 28 Jul. 1903; J.C. Crawford leg.; USNM M.G.R. Database No. 5893 • 1 ♀; same data as for preceding; 16 Aug. 1903; J.C. Crawford leg.; USNM Crawford no. 2268/M.G.R. Database No. 5894. – North Carolina • 1 ♀; North Carolina; USNM M.G.R. Database No. 1339 • 2 ♀♀; Edgecombe County; 28 Aug. 1996; W.A. Mangum leg.; KUNHM M.G.R. Database Nos 1340, 1341 • 1 ♂; same data as for preceding; 29 Aug. 1996; W.A. Mangum leg.; KUNHM M.G.R. Database No. 4596 • 1 ♀; Tuckasegee; 31 Aug. 1957; L.A. Kelton leg.; CNC. – Oklahoma • 1 ♂; Ardmore; 1 Jun. 1909; F.C. Bishopp leg.; USNM M.G.R. Database No. 5590 • 1 ♀; Keystone Wildlife Management Area (Arkansas River), Osage County; 36.3066° N, 96.5221° W; 1 Jul. 2022; M.E. Powley leg.; OKBS OKBS.POL.4924 • 1 ♀; Lake Texoma (2 mi E of Willis); Jun. 1965; R.M. Bohart leg.; UCBME M.G.R. Database No. 11682 • 1 ♂; Sequoyah National Wildlife Refuge, Sequoyah County; 35.4448° N, 95.0120° W; 5 Jul. 2022; M.E. Powley leg.; OKBS OKBS.POL.4958 • 1 ♀; Sequoyah National Wildlife Refuge, Sequoyah County; 35.4500° N, 95.0110° W; 26 Jun. 2023; S. O’Dell leg.; OKBS OKBS.POL.9870. – South Carolina • 1 ♂; Swansea; 8 Aug. 1911; F.K. Knab leg.; USNM M.G.R. Database No. 1410. – Texas • 5 ♂♂; 1877; J. Tosquinet leg.; IRSNB M.G.R. Database Nos 762 to 766 • 2 ♀♀; 16 mi E of Goliad; 7 May 1953; R.H. Beamer leg.; KUNHM M.G.R. Database Nos 4609, 4611 • 1 ♂; same data as for preceding; 7 May 1953; R.H. Beamer leg.; KUNHM M.G.R. Database No. 4594 • 1 ♀; Athens; 6 Aug. 1906; F.C. Bishopp leg.; USNM M.G.R. Database No. 1336 • 1 ♂; Bastrop; 11 May 1954; R.H. Beamer leg.; KUNHM M.G.R. Database No. 4595 • 1 ♂; Colorado County; 12 May 1922; G.O. Wiley leg.; KUNHM M.G.R. Database No. 4593 • 2 ♀♀; Giddings; 9 May 1954; R.H. Beamer leg.; KUNHM M.G.R. Database Nos 1423, 4610 • 1 ♀; Lytle, Atascosa County; 1 Jun. 1951; H.E. Evans leg.; KUNHM M.G.R. Database No. 4613 • 1 ♀; New Boston; 5 Jun. 1909; F.C. Bishopp leg.; USNM M.G.R. Database No. 1338 • 1 ♀; Paris; 5 Aug. 1904; F.C. Bishopp leg.; USNM M.G.R. Database No. 1335 • 1 ♀; Willis; 19