Claspettomyia carpatica Mamaev 1998

Claspettomyia carpatica Mamaev, 1998 Fig. 16A I studied the holotype male of C. carpatica in 2012. A sketch of the terminalia made at that time puts me in a position to conclude that a single Claspettomyia male studied here from Baden-Württemberg belongs to the same species. The specimen from Germany is illustrated here (Fig. 16A). Revised diagnosis Claspettomyia carpatica differs from congeneric species in characters of the male terminalia, as follows. The apical bulge of the gonostylus is moderately large, subglobular, and covered with long dense microtrichia, fine setae, and about 2 stiff bristles (↓ 1, Fig. 16A); the apices of the gonocoxal processes have 2‒3 small knobs subterminally and are bent ventrad (↓ 2); the protuberances near the gonostylar bases are moderately large, pointed, and densely covered with microtrichia (↓ 3); and the parameres are tusk-shaped, moderately long, and slighly bent dorsad (↓ 4). The specimen studied here (ZSM-DIP-42305-D11) corresponds with the holotype in that the eye bridge is 4‒5 ommatidia long dorsally and the neck of the fourth flagellomere is 1.5× as long as the node. Differential diagnosis Claspettomyia carpatica is one of several species resembling C. niveitarsis, a rather common and widespread species in Europe. In contrast to C. carpatica, the gonostylar bulge of C. niveitarsis has 4‒5 stiff bristles, the apices of the gonocoxal processes are smooth, and the neck of the fourth flagellomere is 1.7‒2.1 × as long as the node (Jaschhof & Jaschhof 2013: 280, fig. 132b). A further broadly similar species is C. rossica, whose holotype I examined in 2012. Studied here were also nine males in the Penttinen collection that proved the occurrence of C. rossica in Finland (Jaschhof et al. 2014). From this it appears that C. rossica differs from C. carpatica in the gonostylar bulge, which is slightly more prominent and equipped with up to 5 stiff bristles; in the parameres, whose knobs tend to be more numerous and slightly larger; in the gonocoxal protuberances, which have setae in addition to microtrichia; in the longer eye bridge, which consists of 7‒8 ommatidia dorsally; and in the longer flagellomeral necks, with the neck-to-node ratio being 1.7‒1.9 in the fourth flagellomere. Material examined GERMANY ‒ Baden-Württemberg • 1 ♂; Malsch, Luderbusch; 48°91′31″ N, 8°33′25″ E; elev. 117 m; 28 Jun.‒5 Jul. 2020; D. Doczkal and K. Grabow leg.; Malaise trap; south-facing hill slope; ZSM-DIP-42305-D11. Distribution Germany (new record); Ukraine (Gagné & Jaschhof 2021). Remarks Two males studied here from Rimpar, Bavaria, deviate slightly from the diagnosis given above for C. carpatica: the basal portion of the gonostylus is slightly thicker; the gonocoxal protuberances next to the gonostylar bases are less prominent and devoid of vestiture; the parameres are possibly more strongly bent; the eye bridge is shorter dorsally (2‒3 ommatidia); the neck of the fourth flagellomere is slightly longer (1.7× the node); and the palpus is shorter than the head height (versus longer than the head height in C. carpatica). The two specimens likely represent a species distinct from C. carpatica, which is labeled C. sp. MJDE 5 in the list at the end of this paper. A DNA barcode was obtained here for C. sp. MJDE5 (BIN BOLD:AER0015) but unfortunately not for C. carpatica.