Updated Synthesis of the Upper Cretaceous Vertebrate Assemblages from the Northern Onshore Sector of the Portuguese West Iberian Margin

Vertebrate remains from the Upper Cretaceous of Portugal have been studied for more than 140 years, beginning with the pioneering work of the Geological Commissions and early vertebrate specialists. The Cenomanian of Portugal occupies a key position in understanding vertebrate assemblages from the south-western European Late Cretaceous archipelago. Recent discoveries in the Cenomanian of the onshore northern sector of the Portuguese West Iberian Margin have revealed two new genera of Squamata and Crocodylomorpha, as well as extended biogeographical ranges for groups such as Obaichthyidae. Although Campanian–Maastrichtian fossils are limited to the northern sector, they provide valuable insights into the composition and evolution of western Iberian vertebrate faunas, including significant remains of small dinosaurs and mammals, and the westernmost European occurrences of Pycnodontiformes, Holostei, Elopiformes, Amphibia, Testudines, Squamata, and Crocodylomorpha. The Upper Cretaceous of the Portuguese onshore northern sector of the West Iberian Margin has yielded six vertebrate holotypes, four lectotypes, and syntypes though much material remains undescribed or in need of revision. Identifying additional fossil sites is essential to further expand the dataset. This contribution integrates historical and current knowledge and highlights key priorities for future research.