Changes in pollen production, pollen heteromorphism and ovule production with increased selfing in <i>Viola arvensis</i>

Unlike floral morphology, pollen and ovule production did not evolve rapidly with increased selfing rates, suggesting a delayed change of the pollen:ovule ratio compared to other traits of the selfing syndrome. The absence of correlation between pollen heteromorphism and selfing rate can be explained by multiple (non-mutually exclusive) factors: a decrease in allo-pollen deposition correlated with the evolution of the selfing syndrome, pollinator declines or the absence of selection in this trait.