A new species of genus Tingiometra from mid-Cretaceous Kachin amber (Heteroptera: Tingidae: Tingiometrinae)

Tingidae are also known as ‘lace bugs’ because of the delicate, lace-like appearance of their pronotum and hemelytra. Currently, lace bugs (Heteroptera: Cimicomorpha: Tingidae) comprise approximately more than 2,613 species are placed in about 318 genera (Schuh & Weirauch, 2020), they are extremely important phytophagous pests from all over the world (Drake & Davis, 1960; Drake & Ruhoff, 1965). Tingidae are classified into three extant subfamilies: Tinginae; Cantacaderinae; Vianaidinae, and one extinct subfamily: Tingiometrinae. To date, 46 genera with 82 species of Tingidae are known in the fossil record (Wang et al., 2021; Kaulfuss & Heiss, 2023; Ross, 2024). The subfamily Tinginae comprises 29 genera and 46 species, Cantacaderinae includes 6 genera and 18 species, Vianaidinae consists of 2 genera and 2 species, Tingiometrinae contains 2 genera and 5 species and 7 genera and 11 species remain unassigned to any subfamily (Schuh et al., 2006; Heiss et al., 2015; Du & Yao, 2018; Popov & Golub, 2019; Golub & Heiss, 2020; Poinar & Vega, 2020; Golub et al., 2021; Wang et al., 2021; Ross, 2024), ranging from the Early Cretaceous to the Miocene. Archetingis ladinica Montagna et al., 2018, the earliest fossil record of the Tingidae is from the Middle Triassic in Switzerland, but its classification remains controversial (Wang et al., 2021).